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Source: Unless otherwise indicated, the information in this Research Project Summary comes from the following paper:
Neumann, David D.; Dickmann, Donald I. 2001. Surface burning in a mature stand of Pinus resinosa and Pinus strobus in Michigan: effects on understory vegetation. International Journal of Wildland Fire. 10: 91-101.Prior to the fires, the average density of red pine was 131 trees/ha, dbh ranged from 16 to 40 cm, and average basal area was 29 m²/ha. The average density of white pine trees was 93/ha, dbh measurements ranged from 20 to 60 cm, and basal area averaged 16 m²/ha. Other prominent woody vegetation in the plantation included red maple (Acer rubrum), sugar maple (A. saccharum), hickory (Carya spp.), black cherry (Prunus serotina), white, red, and black oaks (Quercus alba, Q. rubra, Q. velutina), common buckthorn (Rhamnus cathartica), and northern bush honeysuckle (Diervilla lonicera). The most common understory species were violet (Viola spp.), spinulose woodfern (Dryopteris carthusiana), broadleaf enchanter's nightshade (Circaea quadrisulcata), and bedstraw (Galium spp.).
PLANT PHENOLOGYPrescription fires were strip headfires that burned on 2 sites, ignited with drip torches. Fire behavior was similar on the once-burned site and the site burned biennially. Strips were approximately 8 m apart on flat areas and a bit closer as slope steepness increased. Flame lengths ranged from 0.2 to 0.8 meter, and less than 10 pines had over 80% of their needles scorched. Almost all understory vegetation was top-killed, but no more than the top 1.5 cm of the needle-leaf litter layer was consumed. The humus layer was left intact, and no downed wood greater than 2.5 cm in diameter was consumed. The recorded Canadian forest fire weather indices indicate low-severity fires. Fire conditions are summarized below:
Fireline weather conditions | Canadian Forest Fire Weather Indices* | ||||||||
Fire date | Temperature (°C) | Relative humidity (%) | Wind (km/h) | FFMC | DMC | DC | ISI | BUI | FWI |
10 May 1991 | 26-29 | 32-40 | <3.2, gusts to 8 | 90.5 | 23 | 76 | 6.5 | 26 | 11.6 |
10 May 1993 | 28-32 | 20-22 | <3.2, gusts to 8 | 92.7 | 33 | 84 | 8.1 | 34 | 15.6 |
Common name | Unburned | Burned once | Burned twice |
eastern white pine | 35% | 32% | 37% |
red pine | 61% | 64% | 61% |
Hardwood species | 4% | 4% | 2% |
While the overstory was unaffected by fire, the pine and hardwood sapling and large seedling** forest layers decreased significantly (p= 0.05) when burned. Unburned plots had almost double the number of saplings and 15 times the number of large seedlings of plots burned once. Two fires completely removed the sapling layer, but some large seedlings persisted. Below are the densities of woody vegetation by size class and burn treatment in 1995:
Burn treatment | Unburned | Burned once | Burned twice |
Total overstory basal area ± s (m²/ha) |
48.8±7.9a* | 46.7±8.4a | 47.5±11.2a |
Saplings (2-5.9 cm dbh) ± s (number/ha) |
16,111±9,010c | 9,277±7,870b | 0±0a |
Large seedlings** (>1 m, ≤1.9 cm dbh) ± s (number/ha) |
3,944±1,860b | 277±960a | 166±1,130a |
There were no saplings on sites burned twice, and most woody species decreased after single fires. Exceptions were common buckthorn and sassafras (Sassafras albidum), which increased on burned plots, and two species, viburnum (Viburnum spp.) and tuliptree (Liriodendron tulipifera), that occurred only on burned plots. The density of boxelder (A. negundo), red oak, and elderberry (Sambucus spp.) was unchanged by the single fire.
The number of dominant seedlings on burned plots was substantially fewer than on unburned plots. Dominant seedlings included red maple, black cherry, and sugar maple. Many large seedlings occurred only on unburned plots. These were American hornbeam (Carpinus caroliniana), northern bush honeysuckle, green ash (Fraxinus pennsylvanica), and common prickly ash (Zanthoxylum americanum). Below are the densities of large woody seedlings** (<1 m, ≤1.9 cm dbh) on burned and unburned plots as of 1995.
Common name | Scientific name | Unburned | Burned once* |
box elder | Acer negundo | 56 | 56 |
red maple | Acer rubrum | 3,611 | 833 |
sugar maple | Acer saccharum | 1,111 | 667 |
American hornbeam | Carpinus caroliniana | 56 | 0 |
hickory | Carya spp. | 722 | 167 |
hackberry | Celtis occidentalis | 111 | 0 |
flowering dogwood | Cornus florida | 56 | 0 |
beaked hazelnut | Corylus cornuta | 167 | 111 |
northern bush honeysuckle | Diervilla lonicera | 1,278 | 0 |
glossy buckthorn | Frangula alnus | 778 | 444 |
white ash | Fraxinus americana | 333 | 111 |
green ash | Fraxinus pennsylvanica | 333 | 0 |
tuliptree | Liriodendron tulipifera | 0 | 222 |
mulberry | Morus spp. | 111 | 56 |
sweet cherry | Prunus avium | 167 | 56 |
black cherry | Prunus serotina | 3,167 | 500 |
white oak | Quercus alba | 667 | 389 |
red oak | Quercus rubra | 667 | 667 |
black oak | Quercus velutina | 333 | 0 |
common buckthorn | Rhamnus cathartica | 944 | 1,722 |
elderberry | Sambucus spp. | 444 | 444 |
sassafras | Sassafras albidum | 556 | 2,389 |
viburnum | Viburnum spp. | 0 | 444 |
grape | Vitis spp. | 389 | 0 |
common pricklyash | Zanthoxylum americanum | 56 | 0 |
Three years following fire, the coverage of small woody seedlings was significantly (p=0.05) greater on singly and biennially burned plots than unburned plots. Hickory, oak (Quercus spp.), common buckthorn, sumac (Rhus spp.), sassafras, poison-ivy (Toxicodendron radicans), Virginia creeper (Parthenocissus quinquefolia), and blackberry (Rubus spp.) all had greater coverage on both singly and biennially burned plots than on unburned plots. Coverage of Virginia creeper and blackberry was more than double on burned plots. Glossy buckthorn (Frangula alnus) was found only on plots burned biennially. Ash seedling coverage remained relatively unchanged by fire. Rose (Rosa spp.) and grape (Vitis spp.) coverage increased on plots burned once but was lower on plots burned twice. Bristly greenbrier (Smilax tamnoides) coverage was substantially lower on plots burned once, but on plots burned twice coverage was nearly the same as on unburned plots. Maple (Acer spp.), northern bush honeysuckle, pine (Pinus spp.), cherry (Prunus spp.), and basswood (Tilia americana) seedling coverage was greatest on unburned plots. The mean percent cover of small (<1 m) woody seedlings** on burned and unburned plots in 1994 is summarized below.
Common name | Scientific name | Unburned | Burned once | Burned twice |
maple | Acer spp. | 2.11 | 1.57 | 0.44 |
hickory | Carya spp. | 0.28 | 0.51 | 0.38 |
northern bush honeysuckle | Diervilla lonicera | 0.64 | 0.4 | 0.4 |
glossy buckthorn | Frangula alnus | 0 | 0 | 0.28 |
ash | Fraxinus spp. | 0.23 | 0.28 | 0.24 |
Virginia creeper | Parthenocissus quinquefolia | 4.88 | 11.98 | 11.11 |
pine | Pinus spp. | 0.73 | 0.23 | 0.32 |
cherry | Prunus spp. | 1.1 | 0.36 | 0.27 |
oak | Quercus spp. | 0.5 | 1.89 | 0.87 |
common buckthorn | Rhamnus cathartica | 0.53 | 1.9 | 1.5 |
sumac | Rhus spp. | 0 | 0.26 | 0.21 |
rose | Rosa spp. | 0 | 0.11 | 0 |
blackberry | Rubus spp. | 2.73 | 12.66 | 16.73 |
elderberry | Sambucus spp. | 0.08 | 0.01 | 0.48 |
sassafras | Sassafras albidum | 2.68 | 4.78 | 4.78 |
bristly greenbrier | Smilax tamnoides | 0.87 | 0.24 | 0.89 |
basswood | Tilia americana | 0.5 | 0.13 | 0 |
eastern poison-ivy | Toxicodendron radicans | 0.74 | 1.44 | 1.34 |
grape | Vitis spp. | 1.35 | 1.82 | 0.43 |
Total* | 20±24.1b | 40.6±33.5a | 40.7±38 a |
Herbaceous coverage was significantly (p= 0.05) less on unburned than both singly and biennially burned plots. The mean coverage of herbaceous species was double that of unburned plots on both the singly and biennially burned plots. Biennially burned plots had a substantial understory that averaged less than 1 m in height. The understory of once-burned plots that had a longer recovery period averaged 1 to 1.5 m in height. Seventeen of the twenty-three forb species had greater or unchanged coverage on both burned plots than on unburned plots. The rest of the forb species had greatest coverage on either the singly or biennially burned plots. American pokeweed (Phytolacca americana) and groundnut (Apios americana) displayed the greatest coverage differences between burned and unburned plots. Both graminoid genera also had greater coverage on the burned plots. The average percent cover of herbaceous species on burned and unburned plots in 1994 is given below [4].
Common name | Scientific name | Unburned | Burned once | Burned twice |
Forbs | ||||
groundnut | Apios americana | 0.29 | 1.62 | 1.09 |
lesser burrdock | Arctium minus | 0 | 1 | 0.07 |
common milkweed | Asclepias syriaca | 0 | 0.83 | 0.11 |
aster | Aster spp. | 0.08 | 0.91 | 0.14 |
broadleaf enchanter's nightshade | Circaea lutetiana ssp. canadensis | 0.57 | 0.29 | 1.09 |
pointedleaf ticktrefoil | Desmodium glutinosum | 0 | 0.18 | 0.4 |
spinulose woodfern | Dryopteris carthusiana | 2.88 | 4.39 | 3.24 |
Virginia strawberry | Fragaria virginiana | 0.03 | 0.2 | 0.12 |
bedstraw | Galium spp. | 0.43 | 0.52 | 0.3 |
orange hawkweed | Hieracium aurantiacum | 0 | 0.33 | 0.07 |
prickly lettuce | Lactuca serriola | 0 | 0.07 | 0.14 |
partridgeberry | Mitchella repens | 0 | 0.33 | 0 |
Clayton's sweetroot | Osmorhiza claytonii | 0.18 | 0.33 | 0.06 |
mountain woodsorrel | Oxalis montana | 0 | 0.06 | 0.22 |
American lopseed | Phryma leptostachya | 0.23 | 0.32 | 0.16 |
American pokeweed | Phytolacca americana | 0.08 | 1.26 | 4.84 |
Canadian clearweed | Pilea pumila | 0.29 | 0.02 | 0.4 |
mayapple | Podophyllum peltatum | 0 | 0.2 | 0.01 |
smooth Solomon's-seal | Polygonatum biflorum | 0.29 | 0.63 | 0.26 |
common sheep sorrel | Rumex acetosella | 0 | 0.10 | 0.13 |
common dandelion | Taraxacum officinale | 0.04 | 0.11 | 0.13 |
trillium | Trillium spp. | 0 | 0.03 | 0.14 |
violet | Viola spp. | 1.74 | 0.67 | 2.69 |
Graminoids | ||||
sedge | Carex spp. | 0.03 | 0.28 | 0.08 |
bluegrass | Poa spp. | 0.03 | 0.37 | 0.17 |
Total* | 7.3±12.4b | 15.4±20a | 16.4±22.4a |
The researchers provide frequency data in the original paper that are not presented here.
FIRE MANAGEMENT IMPLICATIONS:1. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905]
2. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; Lewis, Mont E.; Smith, Dixie R. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998]
3. Kuchler, A. W. 1964. United States [Potential natural vegetation of the conterminous United States]. Special Publication No. 36. New York: American Geographical Society. 1:3,168,000; colored. [3455]
4. Neumann, David D.; Dickmann, Donald I. 2001. Surface burning in a mature stand of Pinus resinosa and Pinus strobus in Michigan: effects on understory vegetation. International Journal of Wildland Fire. 10: 91-101. [40201]