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| Canyon live oak (tree form). Henry W. Coe State Park, Morgan Hill, California. Photo by Akira. | Canyon live oak (shrub form). Huckleberry Botanic Regional Preserve, Contra Costa County, California. Photo by Jane Huber/Bay Area Hiker. |
Mixed evergreen: Canyon live oak is a dominant subcanopy species in mixed-evergreen forests in the mountains of southern Oregon and California [20,21,25,42,67,79,85,130,191,194]. A canyon live oak phase of mixed-evergreen forest occurs in shady canyons and on steep, north-facing slopes [79]. For information on tree, shrub, and herbaceous species associated with canyon live oak in mixed-evergreen forests, see these sources: [4,8,36,63,168].
Mixed conifer: Canyon live oak is a hardwood associate in mixed-conifer forests in southwest Oregon, California, and northern Baja California [148,153,154,234]. For information on species associated with canyon live oak in mixed-conifer forests, see these sources: [60,153,164,219,220]. Canyon live oak is also a subcanopy component of giant sequoia (Sequoiadendron giganteum) forests in the Sierra Nevada [108,167,237] and upland redwood (Sequoia sempervirens) forests in northern coastal California [97,193].
Bigcone Douglas-fir-canyon live oak: Canyon live oak is codominant with bigcone Douglas-fir (Pseudotsuga macrocarpa) on mesic, steep, north-facing slopes and in ravines in southern California, often within a matrix of chaparral vegetation [20,36,43,90,94,139,141,148,219].
Mixed hardwood and oak woodland: Canyon live oak is an important component of mixed hardwood forests and oak (Quercus spp.) woodlands throughout California [82,92,99,196,219,244]. It grows in California black oak (Q. kelloggii) woodlands in southwestern Oregon and California [42,43,63,99,234,252]. Canyon live oak grows with scrub oak (Q. dumosa) and island oak (Q. tomentella) on Santa Catalina Island [165] and is a component of blue oak (Q. douglasii) woodlands in California foothills [18,20,21,40,162,168]. It is also associated with Oregon white oak (Q. garryana) [99,202], interior live oak (Q. wislizeni), coast live oak (Q. agrifolia), and valley oak (Q. lobata) [43,59,82,90,168].
Chaparral: Canyon live oak is common in montane and mixed-chaparral communities in the foothills and mountains of southwestern Oregon, California, and northern Baja California [19,63,64,75,110,132,152,219,226,227]. It also occurs in chaparral communities in Arizona [47,219].
Other: Canyon live oak is also commonly associated with Coulter pine (Pinus coulteri) on xeric sites within or adjacent to chaparral in central and southern California [20,62,153,219,248] and northern Baja California [150]. Canyon live oak is associated with cypress (Cupressus spp.) groves throughout California [219] and with bristlecone fir (Abies bracteata) forests in the Santa Lucia Mountains of Monterey County, California [91,196,214]. It is common in the understory of singleleaf pinyon (Pinus monophylla) woodlands in southern California [152,228,232,233] and Parry pinyon (P. quadrifolia) forests in northern Baja California [150].
Canyon live oak is listed as a dominant species in the following vegetation classifications:
United States― |
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| Canyon live oak in bloom. Skyline Ridge Open Space Preserve, Palo Alto, California. Photo by Akira. |
Canyon live oak is one of the most morphologically variable oaks in North America [61]. It is a spreading, perennial, sclerophyllous evergreen [61,88,100] that ranges from less than 15 feet to 100 feet (5-30 m) tall and up to 10.7 feet (3.3 m) DBH [88,101,168,179]. Its growth form varies depending on the site [34]. It grows as a shrub and may form dense thickets on mountain slopes and ridgetops, and it grows as a tree in sheltered, moist canyons [43,52,133,150,219]. Its size generally increases with soil depth [161]. In open areas the crown is dense, wide-spreading, and reaches nearly to the ground. In closed stands the crown is smaller in diameter and concentrated in the top half of the tree [179].
Canyon live oak is a thin-barked tree for its size [144,148]. The bark is smooth to flaky. It is fissured in small stems and more deeply furrowed in large stems [88,143,179]. Twigs are 1 to 2 mm in diameter. They are densely pubescent when young and become smooth with age [61,143]. Leaves are thick, leathery, oblong and 0.5 to 4.0 inch (1.3-10.0 cm) long. Leaf margins are usually spiny in young trees and smooth in old trees, although both leaf types are often present on the same tree [6,88,101,143,179]. Canyon live oak begins to produce flowers at approximately 20 years of age [219]. Male flowers are in catkins 2 to 4 inches (5-10 cm) long. Female flowers are solitary or in sparsely-flowered spikes [37,117,219]. Acorns are ellipsoidal, 1 to 3 inches (2.5-7.6 cm) long, and 0.5 to 1.3 inches (1.3-3.3 cm) wide. They are 1-seeded, or rarely 2-seeded, and occur singly or in clusters of 2 to 5 [37,61,88,143,179,219].
Canyon live oak has a deep and extensive root system [89,93,219]. Road cuts have exposed roots up to 24 feet (7.3 m) deep [87]. In coarse soils canyon live oak has a pronounced taproot, whereas in rocky areas the roots are shallow and spreading [219]. Canyon live oak supports ectomycorrhizal fungi that enhance growth of Douglas-fir seedlings in southwest Oregon and northern California forests [7,38]. Canyon live oak is long lived: it may survive for up to 300 years [93,141,168,179,219].
RAUNKIAER [182] LIFE FORM:Pollination: Canyon live oak is wind pollinated [118,219].
Breeding system: Canyon live oak is monoecious [37,219]. Male and female flowers are scattered throughout the crown, which helps to prevent self-fertilization [219].
Seed production: The minimum seed-bearing age in canyon live oak is 20 years [37]. Canyon live oak is a "prolific" seed producer at irregular intervals [179]. Some trees apparently produce acorns every year [219]. Acorn production was highest in canyon live oak out of 5 oak species studied on the Hastings Natural History Reservation in central coastal California. Canyon live oak produced consistently good acorn crops over the 7-year study period [46].
Canyon live oak acorn production varies among years, populations, and individual trees [46,115]. Open-grown trees can produce large acorn crops: up to 400 pounds (180 kg) of acorns per tree have been reported [249]. There are approximately 150 acorns/lb [155], although many acorns are not viable [141]. Trees in sprout clumps tend to produce fewer acorns than larger, single-stemmed trees [219]. In a study of acorn production in the Central Coast Ranges, variation in weather parameters explained 63% of variation in canyon live oak acorn production (P<0.01). High amounts of rainfall had a positive influence in both years of acorn development, while cool temperatures during the first year of acorn development reduced acorn production [116,117]. Mast fruiting may increase pollination and/or satiate acorn predators [118]. Canyon live oak masts approximately every 2 to 6 years [113,219].
Seed dispersal: Canyon live oak acorns are large and usually fall a short distance from the tree [141]. On steep canyon slopes, acorns may roll downhill a long distance [219]. Small mammals and birds that cache canyon live oak acorns are also effective dispersal agents [118,219].
Seed banking: Seed banking in canyon live oak is unlikely because the acorns, lacking a dormant phase, often germinate soon after falling from the tree [37,107]. Some acorns, however, are likely stored in the soil by acorn-caching animals.
Germination: Germination in canyon live oak starts later in the growing season and continues longer than associated oak species [137]. Germination is hypogeal. Canyon live oak acorns germinate best in moist soil covered with leaf litter. Few uncovered acorns germinate [219]. Under greenhouse conditions, canyon live oak acorns planted in peat and loam in deep boxes and covered with 1 inch (2.5 cm) of "firmed" soil germinated in about 60 days [120]. Acorns can germinate in dense shade [219].
Germination rates in canyon live oak vary from 5% to 75% [120,142,155,206]. In a greenhouse experiment, 35% germination occurred after 2 years of storage [177]. In a separate experiment, 50% germination occurred after 35 days at 55 to 68 °F (13-20 °C) with no stratification [137]. Canyon live oak acorns germinate well at temperatures ranging from 41 to 48 °F (5-9 °C) [137]. Cold stratification enhances germination [178,219], although some acorns germinate readily without treatment [104,136]. Canyon live oak may germinate successfully from planted acorns without added water [206].
Seedling establishment/growth: Seedling growth is relatively slow in canyon live oak [141,215]. Seedlings grow best in shade [141,219]. Seedling abundance is high in "favorable years" [93]. Seedlings can be very dense and evenly distributed in canyon live oak stands [219]. In California, there is concern that some oak species are declining due to lack of regeneration. Seedling and sapling density data, however, suggest that canyon live oak does not have a regeneration problem in juvenile size classes. Seedlings were found on 94% of the canyon live oak vegetation type sampled throughout California. Saplings were found on 81% of this type [34]. In the southern Sierra Nevada, canyon live oak seedlings were present in 75% of oak woodland plots sampled, and saplings were present in 48% of plots [200]. In a bigcone Douglas-fir-canyon live oak community in the San Bernardino Mountains, California, canyon live oak density averaged 365 seedlings/ha [141].
Limited data suggest that seedling survival may be high in canyon live oak. In an experiment in the southern California Coast Ranges, 16 germinating canyon live oak acorns were planted in cleared plots. Seedling survival was 94% after 1 year and 69% after 2 years [78]. Recruitment of pole-sized trees may be poor, however. A size-class analysis of canyon live oak in the San Jacinto Mountains of southern California showed high seedling abundance relative to the number of older trees [190]. This difference may or may not signify poor recruitment of canyon live oak into older age classes on some sites. Further research on canyon live oak recruitment beyond the sapling stage is needed.
Vegetative regeneration: Canyon live oak sprouts from the bole or root crown after fire [47,133,141,144,172,198] or other disturbances [53,89,183,195,219]. In recently disturbed stands, sprouting results in a high density of small-diameter (1.1- to 14-inch (2.8-35 cm)) canyon live oak stems [141]. Seedlings may sprout following top-killing disturbance [133]. Sprouts may reach 1 to 3 feet (0.5-1 m) during the first year of growth. Young, "vigorous" plants sprout readily, although large trees produce more and taller sprouts than small trees. Over time, growth is concentrated in a few dominant sprouts, and the total number of sprouts per clump decreases [219].
Canyon live oak sprouts after harvesting but may not require disturbance to sprout. On Skinner Ridge in the San Bernardino Mountains, canyon live oak sprouted in both thinned and control plots. Most trees that showed a strong root crown sprouting response showed a weak bole sprouting response, and vice versa. Root crown sprouting was generally greater than bole sprouting, as shown in the table below [172].
| Sprouting response of canyon live oak in thinned and control plots, San Bernardino Mountains, California [172] | |||
| Plots thinned in 1984 | Plots thinned in 1985 | Control plots | |
| Prethin stems/ha | 3,144 | 2,919 | 3,247 |
| Postthin residual stems/ha | 1,341 | 1,742 | 3,247 |
| Percent sprouting | 62% | 67% | 53% |
| Mean bole sprouts/tree | 21.45 | 24.47 | 23.61 |
| Mean root crown sprouts/tree | 88.53 | 39.52 | 28.89 |
In the eastern Siskiyou Mountains of southwest Oregon, canyon live oak produced more sprouts after clearcutting than after thinning. Sprouts emerged within 3 weeks of cutting. Canyon live oak produced up to 100 sprouts/root crown: more than either tanoak or Pacific madrone. After the first year, canyon live oak sprouts formed dense, overlapping clumps averaging 3.3 feet (1 m) in height [89].
| Sprouting response (stems/ha) in canyon live oak after clearcutting and thinning in the eastern Siskiyou Mountains, Oregon [89] | |||
| Clearcut | Thinned | Control | |
| Pretreatment | 24,092 | 22,486 | 29,405 |
| Immediately after treatment | 0 | 19,768 | 29,405 |
| Posttreatment year 1 | 553,009 | 157,957 | 33,359 |
SITE CHARACTERISTICS:
Canyon live oak occurs on a wide variety of sites ranging from canyon bottoms to
ridgetops [6,133]. It grows under more variable conditions than any other oak in
California [6]. It is commonly found on steep, rocky,
exposed ridges, rock crevices, and canyon slopes [43,81,133,140,148,179].
Canyon live oak also grows in riparian areas [180,229,241,247], sheltered coves,
and deep, moist, shady ravines and canyons [23,27,43,79,88,101,179]. It forms extensive, often pure stands on many of these
sites [91,133,148].
Elevation: Canyon live oak is most common at middle elevations [161,196,249], but can occur between 300 and 9,000 feet (90-2,700 m) across its range [41,61,88,101,103,133,219,245].
| Elevational ranges of canyon live oak | |
| Location | Elevation |
| Arizona | 5,500-7,000 feet (1,700-2,100 m) [103] |
| California | 300-9,000 feet (90-2,700 m) [41,88,133,219] |
| Nevada | 5,500-6,600 feet (1,700-2,000 m) [101] |
| Oregon | 1,600-5,000 feet (488-1,525 m) [219] |
| Sonora and Baja California, Mexico | <6,600 feet (2,000 m) [245] |
Soils: Canyon live oak grows in a wide variety of soils [168,219]. It reaches its greatest size in deep, rich soils in canyon bottoms [168,179]. It also grows on rocky, shallow, infertile soils [112,148,219] but assumes a small, often shrubby growth form on these sites [133,168]. Canyon live oak grows on sedimentary, metasedimentary, and granitic parent materials [219]. It has been documented on serpentine soils in California [230,240], although it may only survive on serpentine if its roots extend into adjacent, nonserpentine soils [86].
Moisture: Moisture conditions vary in communities where canyon live oak is present. Canyon live oak grows in locations that receive as little as 6 inches (150 mm) and as much as 110 inches (2,790 mm) of annual precipitation [219]. Canyon live oak is well adapted to arid conditions [168,188]. It has deep roots that access deep sources of water during summer drought [89]. Its leathery, evergreen leaves have thick epidermal walls and a waxy cuticle that minimize water loss [98,168]. Knops and Koenig [111] describe canyon live oak as a "drought evader" that limits its growth during the dry season.
Temperature: Mean temperatures in the northern portion of canyon live oak's range are 68 to 74 °F (20-23 °C) in the summer and 37 to 41 °F (3-5 °C) in the winter. In the southern portion of its range, average temperatures are 70 to 77 °F (21-25 °C) in the summer and 41 to 45 °F (5-7 °C) in the winter. The frost-free period ranges from 160 to 230 days across its range [219].
SUCCESSIONAL STATUS:Canyon live oak is also an important component of many early seral communities, primarily because of rapid, prolific sprouting that enables it to grow much faster after disturbance than conifers that reproduce only by seed [28,168]. Canyon live oak is an early-successional tree or shrub on favorable sites in mixed-evergreen forests in the northern portion of its range [219]. It is a common pioneer species in thinned and clearcut forests in Oregon [38], northwestern California [134], and southern California [53]. On the Klamath National Forest in northern California, canyon live oak increased in abundance from the interior to the edge of burned and salvaged forest stands [85]. In some areas, vigorous, shade-tolerant sprouters such as canyon live oak slow the development of coniferous forest by interfering with conifer regeneration [63,89].
The ability of canyon live oak to assume multiple growth forms allows it to dominate early-seral, late-seral, and climax stages in some communities. Bigcone Douglas-fir-canyon live oak forests in southern California, for example, are converted to canyon live oak woodland under a regime of frequent fires. In the absence of fire, bigcone Douglas-fir slowly grows above canyon live oak, although canyon live oak remains a community dominant (review by [139]), [141].
SEASONAL DEVELOPMENT:Fire regimes: Canyon live oak grows in a wide variety of plant communities, all of which are subject to periodic or frequent fire [53]. Information below describes fire regimes in plant communities where canyon live oak is most common.
Canyon live oak forest and/or woodland: Pure stands of canyon live oak generally occur within a major vegetation type (e.g., mixed-evergreen forest, mixed-conifer forest, chaparral), each of which has a distinct relationship with fire. Canyon live oak forests and woodlands, therefore, are highly influenced by the fire regimes of surrounding communities. Historic fire regimes in canyon live oak forests and woodlands were characterized by relatively frequent, low- to moderate-severity surface fires with short- to medium-length fire-return intervals [197,199]. Fires occurred mainly in the summer and fall and were medium to large in size [199]. Arno [12] and Paysen and others [169] characterize the fire regime in the Society of American Foresters canyon live oak cover type [133] as mixed severity with a frequency of <35 years [12,169]. Skinner and others [199], however, indicate that historic fire regime reconstruction in canyon live oak forests is problematic due to decay around fire scars. Fire regime information, therefore, is often based on tree ring analyses of surrounding conifer forest communities where fire scars are preserved [199].
Canyon live oak stands with high stem density and abundant ground fuels are susceptible to wildfire [171]. Canyon live oak commonly grows, however, on steep, xeric, rocky sites with little understory vegetation. Fires are relatively infrequent on these sites due to sparse and discontinuous surface fuels [168,169,180,197,199,216]. On very steep slopes, fire is carried by the fuels that accumulate in draws and on the upslope side of canyon live oak trees [199]. Griffin [80] indicates that when they are dense, canyon live oak crowns can carry a fire even when there is little flammable litter on the ground. Stands of large canyon live oaks in deep, moist canyons are somewhat protected from fire. Low-severity surface fires are typical on these sites [80,199].
Mixed-evergreen forest: Mixed-evergreen forests of southern Oregon and northwestern California, where canyon live oak is an important subcanopy species, historically experienced mixed-severity fires [1,209,216]. Sources of ignition included lightning and Native Americans [25]. Mean historic fire-return intervals in mixed-evergreen forest clustered around 15 to 35 years but ranged from 3 to more than 70 years [2,3,131,199,209,216,246]. Mixed-evergreen forests of the Siskiyou and Klamath mountains are drier than those on the coast, with historically shorter fire-return intervals. Fires were of low and mixed severity, with fire-return intervals getting longer with increasing elevation [2,128]. Atzet and Wheeler [13] estimate a 20-year fire-return interval for the Siskiyou Mountains of Oregon. Size and severity of presettlement fires varied across time at the landscape level (review by [96]). Relatively frequent fires prevented fuels from accumulating, thus moderating fire severity in many cases [48]. Surface fires were more common than crown fires [209]. The waxy leaves of canyon live oak and other sclerophylls in the subcanopy, however, fueled fires that were sometimes intense and carried into the conifer overstory. Severely-burned patches of land were generally smaller than surrounding patches of underburned land (review by [1]).
Contemporary fires in mixed-evergreen forests are larger, with more total area experiencing severe fire, compared to historic fire sizes and severities [209,242]. Mean fire-return intervals are longer than intervals before fire exclusion [209,216]. In a fire history study on the Klamath National Forest, Wills and Stuart [246] found no significant differences in presettlement fire-return intervals among 3 Douglas-fir/tanoak-Pacific madrone sites; however, mean fire-return interval of the fire exclusion period was significantly different from those of the presettlement and settlement periods, with intervals between fires increasing greatly around the turn of the twentieth century (P=0.010).
| Means and ranges of fire-return intervals on 3 sites on the Klamath National Forest, California [246] | |||
| Location | Time interval | Mean (years) | Range (years) |
| Site 1 | |||
| Presettlement | 1745-1849 | 17.3 | 5-41 |
| Settlement | 1849-1894 | 15.0 | 8-26 |
| Fire exclusion | 1894-1987 | 46.5 | 43-50 |
| Site 2 | |||
| Presettlement | 1742-1855 | 10.3 | 5-18 |
| Settlement | 1855-1901 | 9.2 | 7-12 |
| Fire exclusion | 1901-1987 | 28.7 | 18-45 |
| Site 3 | |||
| Presettlement | 1752-1849 | 13.9 | 7-25 |
| Settlement | 1849-1913 | 16.0 | 5-25 |
| Fire exclusion | 1913-1987 | 37.0 | 3-71 |
Mixed-conifer forest: Historic ignition sources in mixed-conifer forests included both lightning and Native Americans [166,167,235,238]. The historic fire regime in mixed-conifer forests was characterized by frequent, low- to moderate-severity surface fires [44,102,152,213,238]. Large, severe fires were infrequent in presettlement mixed-conifer forests of Oregon and California [109,153], although crown fires affecting small areas were probably common (review by [238]). In the lower montane zone of the Sierra Nevada, patchy, stand-replacement fires were most common on north-facing slopes and during extended droughts [226]. Historic fire-return intervals in mixed-conifer forests ranged from approximately 3 to 30 years [45,102,131,163,213,226,235]. In giant sequoia groves in the Sierra Nevada, Swetnam and others [211] reported mean fire-return intervals of 5 to 10 years and a maximum fire-return interval of 20 years. Throughout the mixed-conifer zone, mean fire-return intervals varied with site: more mesic and/or sheltered sites burned less often than more xeric and/or exposed sites. Frequent fires prevented accumulation of surface fuels and maintained an open, park-like forest structure (review by [238]).
As a result of fire exclusion since the early 1900s, fires in mixed-conifer forests are now less frequent, larger, and more severe [44,60,153,167,197,238]. The large size and high severity of modern-day fires is related, in part, to greater fuel accumulation during longer fire-free intervals compared to presettlement times. Because small fires are generally suppressed, very large fires are more likely to occur during severe fire weather, such as Santa Ana Winds and heat waves [145]. Canyon live oak mortality is likely to be greatest on steep slopes where dense stands of canyon live oak often occur [152].
Many mixed-conifer forests in the Sierra San Pedro Mártir in northern Baja California still experience an unmanaged fire regime. The fire regime there is characterized by moderate- to high-severity surface fires that create open, park-like stands of mature trees. Fires may be as large as 12,000 acres (5,000 ha) with a relatively long (~50 year) fire-return interval that results from the gradual buildup of litter and shrub cover. Lightning strikes are the primary source of ignition. Fire severity is often high in stands on steep slopes, where canyon live oak is likely to grow, and in stands with a dense subcanopy of canyon live oak and greenleaf manzanita [145,154].
Bigcone Douglas-fir-canyon live oak forest: Bigcone Douglas-fir-canyon live oak forests generally occur on steep, rocky slopes and in moist ravines where fire is infrequent [20,152]. Historically, low-severity fires occurred in high-elevation bigcone Douglas-fir-canyon live oak forests approximately once every decade, while low-elevation forests burned 1 or 2 times each century [105]. These forests, however, often occur within large expanses of chaparral and are therefore also influenced by chaparral fire regimes [153,203]. Canyon live oak can act as a buffer against the spread of severe chaparral fires due to the nonflammable nature of tree-sized canyon live oaks (relative to adjacent chaparral vegetation) [151,152] and the limited accumulation of understory fuels in dense canyon live oak stands [139]. Although stand-replacement fires are rare in bigcone Douglas-fir-canyon live oak forests [153], strong Santa Ana Winds may drive severe chaparral fires into bigcone Douglas-fir-canyon live oak stands, resulting in stand conversion to canyon live oak woodland [139,141,152]. Large, stand-replacement fires have become more common in bigcone Douglas-fir-canyon live oak forests since fire exclusion [153,203]. Bigcone Douglas-fir can sprout from the bole after a moderate-severity surface fire [139]. After a stand-replacement fire, it commonly takes 40 to 70 years for bigcone Douglas-fir to grow through a 16- to 30- foot (5-9 m) canyon live oak canopy [141].
Oak woodland: Throughout its range, canyon live oak is a component of a variety of oak woodland communities. Historically, these communities were maintained by very frequent, low-severity surface fires [22,92,138,199,226]. Fire-return intervals around 8 to 10 years have been reported for both California black oak and blue oak forests and woodlands [22,138]. A century of fire exclusion has led to encroachment by conifers, reducing the extent of many oak woodlands [22,92,167].
Chaparral: Fire regimes in California chaparral communities are variable [106]. Stand flammability and fire severity generally increase with time since burning due to the buildup of shrub cover and dead fuels [131,149]. Most chaparral becomes extremely flammable within 30 to 60 years after the previous fire [146]. Fires in mature chaparral are generally high-severity, fast-moving crown fires [19,41,131,231] that can exceed 30,000 Btus/second/foot and flame lengths of over 50 feet (15 m) [131]. Stand-replacement crown fires are common, especially under dry, windy conditions. Shrub fuels burn readily once ignited [169]. In a typical chaparral fire, all aboveground biomass is consumed except for the largest stems due to horizontal and vertical continuity of stands (review by [149]).
Most estimates of historic fire-return intervals in California chaparral range from 20 to 60 years [41,90,131,250], although fire-return intervals at high elevations and on northerly aspects range from 50 to 100 years [169]. In montane chaparral in the northern Sierra Nevada, the mean fire-return interval was 28 years prior to fire exclusion in the late 1800s and early 1900s, with a range of 16 to 60 years [50,158]. High-severity fires were typical, with most shrubs being top-killed [50].
Some argue that large chaparral fires are an artifact of decades of fire exclusion, which has resulted in a heavy buildup of fuels [27,55,149]. In a review, however, Keeley [105] states that historically, the majority of the landscape in southern California burned in large, Santa Ana Wind-driven fires that occurred 1 to several times each century. Fire frequency in southern California chaparral was historically greatest in the summer due to dry fuels and the high frequency of lightning storms. These fires were relatively small and may have acted as fuel breaks against larger fires. This mixed-severity fire regime perpetuated a patchy mosaic of age classes in chaparral stands [41,152]. In his review, Keeley states that fire suppression has not effectively excluded fire from southern California chaparral and, although fire size has decreased due to habitat fragmentation, large fires are a natural part of the chaparral fire regime [105].
The following table provides fire regime information that may be relevant to canyon live oak. Find further fire regime information for the plant communities in which this species may occur by entering the species name in the FEIS home page under "Find Fire Regimes".
| Fire regime information on vegetation communities in which canyon live oak may occur. For each community, fire regime characteristics are taken from the LANDFIRE Rapid Assessment Vegetation Models [123]. These vegetation models were developed by local experts using available literature, local data, and/or expert opinion as documented in the .PDF file linked from the name of each Potential Natural Vegetation Group listed below. Cells are blank where information is not available in the Rapid Assessment Vegetation Model. | ||||||||||
| ||||||||||
| Pacific Northwest | ||||||||||
| Vegetation Community (Potential Natural Vegetation Group) | Fire severity* | Fire regime characteristics | ||||||||
| Percent of fires | Mean interval (years) |
Minimum interval (years) |
Maximum interval (years) |
|||||||
| Northwest Woodland | ||||||||||
| Pine savannah (ultramafic) | Replacement | 7% | 200 | 100 | 300 | |||||
| Surface or low | 93% | 15 | 10 | 20 | ||||||
| Oregon white oak | Replacement | 3% | 275 | |||||||
| Mixed | 19% | 50 | ||||||||
| Surface or low | 78% | 12.5 | ||||||||
| Northwest Forested | ||||||||||
| Oregon coastal tanoak | Replacement | 10% | 250 | |||||||
| Mixed | 90% | 28 | 15 | 40 | ||||||
| Mixed conifer (southwestern Oregon) | Replacement | 4% | 400 | |||||||
| Mixed | 29% | 50 | ||||||||
| Surface or low | 67% | 22 | ||||||||
| California mixed evergreen (northern California) | Replacement | 6% | 150 | 100 | 200 | |||||
| Mixed | 29% | 33 | 15 | 50 | ||||||
| Surface or low | 64% | 15 | 5 | 30 | ||||||
| Red fir | Replacement | 20% | 400 | 150 | 400 | |||||
| Mixed | 80% | 100 | 80 | 130 | ||||||
| California | ||||||||||
| Vegetation Community (Potential Natural Vegetation Group) | Fire severity* | Fire regime characteristics | ||||||||
| Percent of fires | Mean interval (years) |
Minimum interval (years) |
Maximum interval (years) |
|||||||
| California Shrubland | ||||||||||
| Chaparral | Replacement | 100% | 50 | 30 | 125 | |||||
| Montane chaparral | Replacement | 34% | 95 | |||||||
| Mixed | 66% | 50 | ||||||||
| California Woodland | ||||||||||
| California oak woodlands | Replacement | 8% | 120 | |||||||
| Mixed | 2% | 500 | ||||||||
| Surface or low | 91% | 10 | ||||||||
| California Forested | ||||||||||
| California mixed evergreen | Replacement | 10% | 140 | 65 | 700 | |||||
| Mixed | 58% | 25 | 10 | 33 | ||||||
| Surface or low | 32% | 45 | 7 | |||||||
| Coast redwood | Replacement | 2% | ≥1,000 | |||||||
| Surface or low | 98% | 20 | ||||||||
| Mixed conifer (North Slopes) | Replacement | 5% | 250 | |||||||
| Mixed | 7% | 200 | ||||||||
| Surface or low | 88% | 15 | 10 | 40 | ||||||
| Mixed conifer (South Slopes) | Replacement | 4% | 200 | |||||||
| Mixed | 16% | 50 | ||||||||
| Surface or low | 80% | 10 | ||||||||
| Jeffrey pine | Replacement | 9% | 250 | |||||||
| Mixed | 17% | 130 | ||||||||
| Surface or low | 74% | 30 | ||||||||
| Mixed evergreen-bigcone Douglas-fir (southern coastal) | Replacement | 29% | 250 | |||||||
| Mixed | 71% | 100 | ||||||||
| Red fir-white fir | Replacement | 13% | 200 | 125 | 500 | |||||
| Mixed | 36% | 70 | ||||||||
| Surface or low | 51% | 50 | 15 | 50 | ||||||
| Southwest | ||||||||||
| Vegetation Community (Potential Natural Vegetation Group) | Fire severity* | Fire regime characteristics | ||||||||
| Percent of fires | Mean interval (years) |
Minimum interval (years) |
Maximum interval (years) |
|||||||
| Southwest Shrubland | ||||||||||
| Interior Arizona chaparral | Replacement | 100% | 125 | 60 | 150 | |||||
| Southwest Woodland | ||||||||||
| Pinyon-juniper (mixed fire regime) | Replacement | 29% | 430 | |||||||
| Mixed | 65% | 192 | ||||||||
| Surface or low | 6% | >1,000 | ||||||||
| Great Basin | ||||||||||
| Vegetation Community (Potential Natural Vegetation Group) | Fire severity* | Fire regime characteristics | ||||||||
| Percent of fires | Mean interval (years) |
Minimum interval (years) |
Maximum interval (years) |
|||||||
| Great Basin Shrubland | ||||||||||
| Interior Arizona chaparral | Replacement | 88% | 46 | 25 | 100 | |||||
| Mixed | 12% | 350 | ||||||||
| Montane chaparral | Replacement | 37% | 93 | |||||||
| Mixed | 63% | 54 | ||||||||
|
*Fire Severities: Replacement=Any fire that causes greater than 75% top removal of a vegetation-fuel type, resulting in general replacement of existing vegetation; may or may not cause a lethal effect on the plants. Surface or low=Any fire that causes less than 25% upper layer replacement and/or removal in a vegetation-fuel class but burns 5% or more of the area. Mixed=Any fire burning more than 5% of an area that does not qualify as a replacement, surface, or low-severity fire; includes mosaic and other fires that are intermediate in effects [84,122]. |
||||||||||
 |
|
| Sprouting canyon live oak after the Cedar Fire in Cuyamaca Rancho State Park. Photo by Linnea Spears. |
IMMEDIATE FIRE EFFECT ON PLANT:
Although canyon live oak is easily top-killed by fires of even relatively low severity [148,179,198,238],
the entire tree is rarely killed [80,93]. Small
canyon live oak trees and shrubs are more vulnerable to fire than large trees [148,176].
Even relatively large canyon live oak trees, however, have thin, flaky outer
bark that is easily ignited [48,171,180]. The bark can ignite even during
a low-severity surface fire, carrying fire up the
trunk and into the crown [80]. Trees can experience fatal
cambium damage when the trunk and/or crown is burned [147,176,183].
Leaves may be heat-killed even if they do not burn [176]. Mortality may also
result from root crown damage caused by smoldering leaf litter at the base of
trees [239]. Canyon live oak acorns are generally destroyed
by fire [93]. Fire triggers a strong sprouting response in
canyon live oak, which is described in detail below.
DISCUSSION AND QUALIFICATION OF FIRE EFFECT:
The probability of fire-induced mortality in canyon live oak increases with increasing height of stem-bark char and
relative char height (height of stem-bark char as a proportion of tree height)
and decreases with increasing DBH and tree height. These parameters were
measured following the Stanislaus Complex Fires in the central
Sierra Nevada in the summer of 1987 [183] and are presented in
the table below.
| Characteristics of live and dead canyon live oak trees following summer wildfire in the central Sierra Nevada [183] | ||||
| DBH range (cm) | Tree height range (m) | Height of stem-bark char (m) | Relative char height | |
| Live trees | 15-51 | 6-16 | 0-1 | 0-0.17 |
| Dead trees | 9-35 | 4-25 | 0.3-15 | 0.03-1 |
Sprouting: Canyon live oak regenerates by sprouting from the root crown or bole after it is damaged or top-killed by fire [47,133,141,144,172,198]. Canyon live oak can sprout within weeks after a fire [168], advancing regeneration and helping to maintain canyon live oak cover [133]. Top-killed canyon live oak exhibit a particularly strong sprouting response. Following the Stanislaus Complex Fires, 100% of top-killed canyon live oak trees sprouted from the root crown, compared to sprouting in 36% of trees that were not top-killed [183]. Within 18 months of the November 1975 Village Fire on the Angeles National Forest, southern California, 90% of top-killed canyon live oak trees had sprouted [176]. Sprouting also occurs when canyon live oak is only slightly damaged by fire [148]. Sprouting may be triggered when sufficient cambial and/or crown damage disrupts the production of growth-regulating hormones that normally inhibit sprouting of dormant buds [183].
Canyon live oak may sprout in the same season it is burned. Within 1 month of the August, 1977 Marble Cone Fire in Monterey County, California, top-killed canyon live oak trees were sprouting "vigorously" from the base of charred trunks. By November of that year, many of the sprouts were several feet tall [80]. Within 3 months of a July, 1960 wildfire on the San Dimas Experimental Forest in southern California, 56% of canyon live oak trees were sprouting. Within 4.5 months, 66% were sprouting [175]. Sprouting response was strongest at low elevation.
| Effect of elevation on sprouting in canyon live oak 4.5 months after wildfire in southern California [175] | |
| Elevation (feet) | % Sprouting |
| 2,100 | 73 |
| 4,100 | 60 |
Seeds and/or seedlings: In some cases, canyon live oak establishes from soil-stored acorns after fire [213]. Because canyon live oak acorns are generally destroyed by fire [93], however, seedling establishment is more likely from off-site seed sources or from crown-stored acorns that survive the fire. The Marble Cone Fire crowned in portions of the summit forest area and changed to a surface fire as it burned downhill into the slope forest area. Following the fire, canyon live oak regenerated from both sprouts and acorns in both forest areas [213].
| Constancy (%) of canyon live oak before and after the 1977 Marble Cone Fire [213] | ||||
| Summit forest (crown fire) | Slope forest (surface fire) | |||
| Prefire | Postfire | Prefire | Postfire | |
| Sprouts | 60 | 100 | 20 | 100 |
| Seedlings | 0 | 100 | 0 | 100 |
Canyon live oak seedling density increased moderately following a low-severity prescribed fire in a Jeffrey pine (Pinus jeffreyi)-California black oak forest in Cuyamaca Rancho State Park, California. Canyon live oak sapling density decreased significantly on burned plots (P<0.02) [135]. For further information, see the Research Project Summary of Martin and Lathrop's [125,135] study.
Repeated burning: Repeated burning favors the shrubby growth form of canyon live oak [148]. Canyon live oak woodland can convert to chaparral with repeated burning. In the absence of fire, the chaparral may revert back to canyon live oak woodland when canyon live oak basal sprouts become tall enough (approximately 15 feet (4.6 m)) to shade out chaparral shrubs [133]. Fire-free intervals of 50 to 100 years may be required for the development of tree-sized canyon live oaks [151]. In general, oaks (Quercus spp.) die if they are burned too frequently [180].
Delayed mortality: The full effect of fire on canyon live oak may not be apparent for several months, or even years, following burning. Heat-injured canyon live oak trees may not die until 8 years after a fire [159]. The Village Fire burned through a dense stand of 6-to 12-inch (15-31 cm) DBH canyon live oak. On most trees, initial damage was apparently restricted to charring at the bottom 1 to 5 feet (0.3-1.5 m) of the trunk and leaf kill of one-third to two-thirds of the crown. Within 18 months, however, all trees 3 inches (7.6 cm) DBH or smaller were top-killed. Mortality from crown damage developed more slowly in larger trees. At 18 months, at least 55% of the larger trees seemed alive, even though most appeared girdled at the base. During the next 18 months, however, approximately 50% of these trees died [176].
| Percent of living canyon live oak crowns 18 and 36 months after fire on the Angeles National Forest, California [176] | |||
| Tree DBH (inches) | 0-3 | 3-6 | 6-12 |
| % live crown after 18 months | 0 | 24 | 74 |
| % live crown after 36 months | 0 | 15 | 46 |
Following prescribed burning in a canyon live oak forest on the San Bernardino National Forest, canyon live oak mortality increased 4% to 5% between postfire years 2 and 6 [171]. For further information on this study, see the Research Project Summary of Narog and others's [159,160,161] study.
DISCUSSION AND QUALIFICATION OF PLANT RESPONSE:| Condition of canyon live oak trees 36 months after fire [176] | |||||
| Tree DBH (inches) | Number of trees |
Tree condition (%) |
|||
| Top-killed | Sprouting | Sprouting + live crown | Live crown only; no sprouting | ||
| 0-3 | 10 | 10 | 90 | 0 | 0 |
| 3-6 | 59 | 3 | 73 | 22 | 2 |
| 6-12 | 97 | 1 | 25 | 70 | 4 |
Fall prescribed fire in a canyon live oak forest on the San Bernardino National Forest resulted in low mortality of canyon live oak >6 inches (15 cm) DBH and high mortality in smaller trees [171]. Some large trees died, however, possibly because of crown scorch or root crown damage caused by long fire residence time in the leaf litter accumulated at the base of trees [170,171,239]. For further information on this study, see the Research Project Summary of the study by Narog and others [159,160,161].
FIRE MANAGEMENT CONSIDERATIONS:Prescribed burning may be used to reduce fire hazard in canyon live oak communities [166,239]. Because canyon live oak trees are easily damaged by fire [148,179,198,238], use of fire alone to remove fuels in canyon live oak communities is generally not a viable management option [77]. Prescribed fire may be a useful tool only in limited situations, such as in a stand of large trees with low fuel loads where trunks are protected from direct heat damage [176]. Thinning prior to burning may mitigate damage to canyon live oak [77,171]. Thinning and prescribed burning in a canyon live oak forest on the San Bernardino National Forest resulted in increased growth in canyon live oak [161] and reduced fire hazard [161,171]. For further information on this study, see the Research Project Summary of Narog and others's [159,160,161] study.
Mortality of canyon live oak trees following burning is often difficult to assess in the first few postfire years. Discolored inner bark and cambium and a fermented smell are indicators of tissue damage [176]. Mechanical assessment of tissue damage using a shigometer may allow for early detection of injury and mortality in canyon live oak, thereby influencing the type of management action taken in burned stands [159]. Plumb [176] provides harvesting recommendations for fire-damaged canyon live oak trees based on tree size and degree of trunk char.Canyon live oak acorns are a primary food source for acorn woodpeckers [82,119]. The acorns are also an important food source for western scrub jays, Steller's jays, band-tailed pigeons, wild turkeys, mountain quail, ground squirrels, western gray squirrels, woodrats, common crows, gophers, American black bears, and mule deer [95,107,115,210,219].
Palatability/nutritional value: Palatability of canyon live oak foliage is generally low [189], although it is considered highly palatable to Columbian black-tailed deer (Orr 1937, cited in [224]). Crude protein content of canyon live oak browse varies from 4.1% to 11.6% depending on the season and location [24,26,126,189,212]. Protein content is higher in canyon live oak leaves than in stems [212]. Fat content ranges from 2.4% to 8.7%, and fiber content ranges from 12.7% to 32.4% [24,126]. Additional nutritional information is provided by Bean and Saubel [24] and Lawrence and Biswell [126].
Canyon live oak acorns are highly palatable to many birds and mammals [95,107,115,219]. In a nutritional study, protein content of canyon live oak acorns was 3.9%, carbohydrate content was 12.6%, lipid content was 16.7%, and energy content was 11.39 kJ/g [114]. American black bears may prefer canyon live oak acorns over interior live oak acorns because canyon live oak acorns have a lower concentration of tannins (Fleck and Layne 1990, cited in [210]).
Cover value: Canyon live oak provides cover for many species of mammals, birds, reptiles, amphibians, and invertebrates. The type and quality of cover provided by canyon live oak is determined in large part by its growth form. Shrubby thickets provide readily available browse and can serve as good hiding or nesting cover for many small birds and mammals. Large trees provide perching, nesting, resting, and foraging sites for numerous species of birds and provide shade and cover for both large and small mammals [189].
Mammals: Canyon live oak forest and woodlands provide habitat for a variety of large mammals including American black bears, deer, coyotes, mountain lions, bobcats, and American badgers [72,161] and small mammals including red tree voles [66], deer mice, white-footed mice, pinyon mice, California pocket mice, California voles, Botta's pocket gophers, and broad-footed mice [32].
Birds: A variety of birds forage in canyon live oak forests and woodlands. In Kern County, California, foraging birds include Nuttall's woodpeckers, acorn woodpeckers, white-breasted nuthatches, plain titmice, black-headed grosbeaks, and Baltimore orioles [29,156]. California spotted owls use a variety of habitats where canyon live oak is common, including canyon live oak forest, mixed-conifer forest, mixed-evergreen forest, riparian hardwood forest, and oak woodlands [33,65,83,161,201,229,238]. In the southern Coast Ranges of California, canyon live oak was the dominant tree at 91% of the sites below 3,600 feet (1,100 m) elevation where California spotted owls were found [70]. On the Sierra National Forest and in Sequoia National Park, California, canyon live oak, interior live oak, and California black oak were most commonly used for California spotted owl nests in foothill riparian and oak woodland habitats. Nesting platforms were found in living canyon live oak trees at a mean height of 47.5 feet (14.5 m). Canyon live oak nest trees were larger than the average canyon live oak tree size: nest trees averaged 18.4 inches (47 cm) in diameter compared to the average diameter of 10.4 inches (26.4 cm). Mean nest tree height was 55 feet (17 m). All of the nests found in canyon live oak trees were platform structures that were likely abandoned squirrel, raptor, or raven nests. The use of platforms allows for California spotted owl reproduction in woodland habitats where trees are not large enough to support cavity nests [201].
Reptiles and amphibians: Hollows and cavities in oak trunks are used for nesting and shelter by a variety of herptiles. Several species of reptiles and amphibians are associated with canyon live oak stands in southern California, particularly on mesic, north-facing slopes. These include San Bernardino Mountain kingsnakes, San Gabriel Mountain slender salamanders, large-blotched salamanders, black-bellied slender salamanders, California slender salamanders, Gilbert's skinks, yellow-blotched salamanders, foothill yellow-legged frogs, western fence lizards, and southern alligator lizards. Many of these are Sensitive Species or Species of Conservation Concern in California [30,31,32,33,204].
Invertebrates: A variety of invertebrates are found in canyon live oak forests [160]. For information on insects found in a canyon live oak forest on the San Bernardino National Forest and the effects of prescribed burning on invertebrate abundance, see the Research Project Summary of Narog and others's [159,160,161] study.
VALUE FOR REHABILITATION OF DISTURBED SITES:Canyon live oak wood is the highest quality of all western oaks [221]. The wood is particularly hard and is sometimes referred to as "rock oak" or "maul oak". It has been used for axles, tool handles, mauls, wagon tongues, plow beams, ship frames, and wheels [124,161,187]. Wedges made from canyon live oak were used to split redwood into railroad ties [127]. Today, its commercial value is limited by the small quantities available [161,221].
Canyon live oak is an attractive landscaping tree [93,120,219] and an important source of firewood in southern California [184,239].
OTHER MANAGEMENT CONSIDERATIONS:General information on diseases in canyon live oak and other California oaks is provided by Raabe [181]. Canyon live oak is less susceptible to the oak wilt pathogen Ceratocystis fagacearum than the deciduous red or black oaks (subgenus Erythrobalanus) in California. In an inoculation experiment, canyon live oak was the least susceptible to oak wilt out of 7 species tested. Where canyon live oak grows in association with California black oak, however, it may be more susceptible to oak wilt [10]. Canyon live oak is a host of Phytophthora ramorum, the pathogen that causes sudden oak death disease in tanoak and many species of oak (Quercus spp.) in California and Oregon [174,185].
Canyon live oak is host to at least 39 cynipid gall wasp species in California [57].1. Agee, James K. 1990. The historical role of fire in Pacific Northwest forests. In: Walstad, John D.; Radosevich, Steven R.; Sandberg, David V., eds. Natural and prescribed fire in Pacific Northwest forests. Corvallis, OR: Oregon State University Press: 25-38. [46954]
2. Agee, James K. 1991. Fire history along an elevational gradient in the Siskiyou Mountains, Oregon. Northwest Science. 65(4): 188-199. [16293]
3. Agee, James K. 1991. Fire history of Douglas-fir forests in the Pacific Northwest. In: Ruggiero, Leonard F.; Aubry, Keith B.; Carey, Andrew B.; Huff, Mark H., technical coordinators. Wildlife and vegetation of unmanaged Douglas-fir forests. Gen. Tech. Rep. PNW-GTR-285. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station: 25-33. [17303]
4. Agee, James K.; Edmonds, Robert L. 1992. Appendix E: Forest protection in the Pacific Northwest. In: Northern Spotted Owl Recovery Team. Final draft--Recovery plan for the northern spotted owl. Vol. 2: appendixes. Portland, OR: U.S. Department of the Interior, Fish and Wildlife Service, Pacific Region: 180-244. [30020]
5. Aizen, Marcelo A.; Patterson, William A., III. 1990. Acorn size and geographical range in the North American oaks (Quercus L.). Journal of Biogeography. 17: 327-332. [13450]
6. Alderman, DeForest C. 1974. Native edible fruits, nuts, vegetables, herbs, spices, and grasses of California: I. Fruit trees and nuts. Leaflet 75-LE/ 2226. Berkeley, CA: University of California, Division of Agricultural Sciences, Cooperative Extension. 14 p. [67651]
7. Amaranthus, M. P.; Perry, D. A. 1989. Interaction effects of vegetation type and Pacific madrone soil inocula on survival, growth, and mycorrhiza formation of Douglas-fir. Canadian Journal of Forest Research. 19: 550-556. [7414]
8. Amaranthus, M.; McNabb, D. H. 1983. Bare soil exposure following logging and prescribed burning in southwest Oregon. In: New forests for a changing world: Proceedings of the 1983 convention of the Society of American Foresters; 1983 October 16-20; Portland, OR. Bethedsa, MD: Society of American Foresters: 234-237. [12362]
9. Anderson, M. Kat; Moratto, Michael J. 1996. Native American land-use practices and ecological impacts. In: Status of the Sierra Nevada. Sierra Nevada Ecosystem Project: Final report to Congress. Volume II: Assessments and scientific basis for management options. Wildland Resources Center Report No. 37. Davis, CA: University of California, Centers for Water and Wildland Resources: 187-206. [28967]
10. Appel, David N. 1994. The potential for a California oak wilt epidemic. Journal of Arboriculture. 20(2): 79-86. [40852]
11. Arizona Fish and Game Department, comp. 1977. The Arizona white-tailed deer. Special Report No. 6. [Federal Aid in Wildlife Restoration Act: Project W-53-R]. Phoenix, AZ: Arizona Fish and Game Department. 108 pp. [61802]
12. Arno, Stephen F. 2000. Fire in western forest ecosystems. In: Brown, James K.; Smith, Jane Kapler, eds. Wildland fire in ecosystems: Effects of fire on flora. Gen. Tech. Rep. RMRS-GTR-42-vol. 2. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 97-120. [36984]
13. Atzet, Thomas; Wheeler, David L. 1982. Historical and ecological perspectives on fire activity in the Klamath Geological Province of the Rogue River and Siskiyou National Forests. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Region. 16 p. [6252]
14. Atzet, Thomas; Wheeler, David L.; Smith, Brad; Franklin, Jerry; Riegel, Gregg; Thornburgh, Dale. 1992. Vegetation. In: Hobbs, Stephen D.; Tesch, Steven D.; Owston, Peyton; Stewart, Ronald; Tappeiner, John C., II; Wells, Gail E., eds. Reforestation practices in southwest Oregon and northern California. Corvallis, OR: Oregon State University, Forest Research Laboratory: 92-113. [69206]
15. Atzet, Thomas; White, Diane E.; McCrimmon, Lisa A.; Martinez, Patricia A.; Fong, Paula Reid; Randall, Vince D., tech. coords. 1996. Field guide to the forested plant associations of southwestern Oregon. Tech. Pap. R6-NR-ECOL-TP-17-96. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Region. Available online: https://www.fs.usda.gov /r6/siskiyou/guide.htm [2004, October 7]. [49881]
16. Atzet, Tom; Wheeler, David; Smith, Brad; Riegel, Gregg; Franklin, Jerry. 1985. The tanoak series of the Siskiyou region of southwest Oregon. Forestry Intensified Research. Corvallis, OR: Oregon State University. 6(4): 6-10. [8594]
17. Baker, Frederick S. 1949. A revised tolerance table. Journal of Forestry. 47: 179-181. [20405]
18. Baker, Gail A.; Rundel, Philip W.; Parsons, David J. 1981. Ecological relationships of Quercus douglasii (Fagaceae) in the foothill zone of Sequoia National Park, California. Madrono. 28(1): 1-12. [6477]
19. Bancroft, Larry. 1979. Fire management plan: Sequoia and Kings Canyon National Parks. San Francisco, CA: U.S. Department of the Interior, National Park Service, Western Region. 190 p. [11887]
20. Barbour, Michael G. 1987. Community ecology and distribution of California hardwood forests and woodlands. In: Plumb, Timothy R.; Pillsbury, Norman H., technical coordinators. Proceedings of the symposium on multiple-use management of California's hardwood resources; 1986 November 12-14; San Luis Obispo, CA. Gen. Tech. Rep. PSW-100. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 18-25. [5356]
21. Barbour, Michael G. 1988. Californian upland forests and woodlands. In: Barbour, Michael G.; Billings, William Dwight, eds. North American terrestrial vegetation. Cambridge; New York: Cambridge University Press: 131-164. [13880]
22. Barnhart, Stephen J.; McBride, Joe R.; Warner, Peter. 1996. Invasion of northern oak woodlands by Pseudotsuga menziesii (Mirb.) Franco in the Sonoma Mountains of California. Madrono. 43(1): 28-45. [51847]
23. Bauer, H. L. 1930. Vegetation of the Tehachapi Mountains, California. Ecology. 11(2): 263-280. [15102]
24. Bean, Lowell John; Saubel, Katherine Siva. 1972. Telmalpakh: Cahuilla Indian knowledge and usage of plants. Banning, CA: Malki Museum. 225 p. [35898]
25. Bingham, Bruce B.; Sawyer, John O., Jr. 1991. Distinctive features and definitions of young, mature, and old-growth Douglas-fir/hardwood forests. In: Ruggiero, Leonard F.; Aubry, Keith B.; Carey, Andrew B.; Huff, Mark H., technical coordinators. Wildlife and vegetation of unmanaged Douglas-fir forests. Gen. Tech. Rep. PNW-GTR-285. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station: 363-377. [17328]
26. Bissell, Harold D.; Strong, Helen. 1955. The crude protein variations in the browse diet of California deer. California Fish and Game. 41(2): 145-155. [10524]
27. Biswell, H. H.; Taber, R. D.; Hedrick, D. W.; Schultz, A. M. 1952. Management of chamise brushlands for game in the north coast region of California. California Fish and Game. 38(4): 453-484. [13673]
28. Biswell, Harold H. 1974. Effects of fire on chaparral. In: Kozlowski, T. T.; Ahlgren, C. E., eds. Fire and ecosystems. New York: Academic Press: 321-364. [14542]
29. Block, William M.; Morrison, Michael L. 1987. Conceptual framework and ecological considerations for the study of birds in oak woodlands. In: Plumb, Timothy R.; Pillsbury, Norman H., technical coordinators. Proceedings of the symposium on multiple-use management of California's hardwood resources; 1986 November 12-14; San Luis Obispo, CA. Gen. Tech. Rep. PSW-100. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 163-173. [5372]
30. Block, William M.; Morrison, Michael L. 1991. Influence of scale on the management of wildlife in California oak woodlands. In: Standiford, Richard B., technical coordinator. Proceedings of the symposium on oak woodlands and hardwood rangeland management; 1990 October 31 - November 2; Davis, CA. Gen. Tech. Rep. PSW-126. Berkeley, CA: Pacific Southwest Research Station: 96-104. [18896]
31. Block, William M.; Morrison, Michael L. 1998. Habitat relationships of amphibians and reptiles in California oak woodlands. Journal of Herpetology. 32(1): 51-60. [52670]
32. Block, William M.; Morrison, Michael L.; Slaymaker, John C.; Jongejan, Gwen. 1988. Design considerations for the study of amphibians, reptiles, and small mammals in California's oak woodlands: temporal and spatial patterns. In: Szaro, Robert C.; Severson, Kieth E.; Patton, David R., technical coordinators. Management of amphibians, reptiles, and small mammals in North America: Proceedings of the symposium; 1988 July 19-21; Flagstaff, AZ. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 247-253. [7115]
33. Block, William M.; Morrison, Michael L.; Verner, Jared. 1990. Wildlife and oak-woodland interdependency. Fremontia. 18: 72-76. [51874]
34. Bolsinger, Charles L. 1988. The hardwoods of California's timberlands, woodlands, and savannas. Resour. Bull. PNW-RB-148. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 148 p. [5291]
35. Bolsinger, Charles L.; Waddell, Karen L. 1993. Area of old-growth forests in California, Oregon, and Washington. Resource Bulletin PNW-RB-197. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 26 p. [24150]
36. Bond, Monica L.; Gutierrez, R. J.; Franklin, Alan B.; LaHaye, William S.; May, Christopher A.; Seamans, Mark E. 2002. Short-term effects of wildfires on spotted owl survival, site fidelity, mate fidelity, and reproductive success. Wildlife Society Bulletin. 30(4): 1022-1028. [44144]
37. Bonner, Franklin T. [In press]. Quercus L.--oak, [Online]. In: Bonner, Franklin T.; Nisley, Rebecca G.; Karrfait, R. P., coords. Woody plant seed manual. Agric. Handbook 727. Washington, DC: U.S. Department of Agriculture, Forest Service (Producer). Available: http://www.nsl.fs.usda.gov/wpsm/Quercus.pdf [2006, July 19]. [62581]
38. Borchers, Susan L.; Perry, David A. 1990. Growth and ectomycorrhiza formation of Douglas-fir seedlings grown in soils collected at different distances from pioneering hardwoods in southwest Oregon clear-cuts. Canadian Journal of Forest Research. 20(6): 712-721. [12989]
39. Borchert, Mark. 1985. Serotiny and cone-habit variation in populations of Pinus coulteri (Pinaceae) in the southern Coast Ranges of California. Madrono. 32(1): 29-48. [5997]
40. Borchert, Mark. 1994. SRM 201: Blue oak woodland. In: Shiflet, Thomas N., ed. Rangeland cover types of the United States. Denver, CO: Society for Range Management: 11. [66660]
41. Botti, Stephen. 1979. Natural, conditional, and prescribed fire management plan. Washington, DC: U.S. Department of the Interior, National Park Service, Yosemite National Park. 51 p. [20901]
42. Bowcutt, Frederica S. 1999. A floristic study of Sugarloaf Ridge State Park, Sonoma County, California. Aliso. 18(1): 19-34. [40636]
43. Boyd, Steve. 1999. Vascular flora of the Liebre Mountains, western Transverse Ranges, California. Aliso. 18(2): 93-139. [40639]
44. Brown, Richard T.; Agee, James K.; Franklin, Jerry F. 2004. Forest restoration and fire: principles in the context of place. Conservation Biology. 18(4): 903-912. [50091]
45. Caprio, Anthony C.; Swetnam, Thomas W. 1995. Historic fire regimes along an elevational gradient on the west slope of the Sierra Nevada, California. In: Brown, James K.; Mutch, Robert W.; Spoon, Charles W.; Wakimoto, Ronald H., tech. coords. Proceedings: symposium on fire in wilderness and park management; 1993 March 30-April 1; Missoula, MT. Gen. Tech. Rep. INT-GTR-320. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station: 173-179. [26217]
46. Carmen, William J.; Koenig, Walter D.; Mumme, Ronald L. 1987. Acorn production by five species of oaks over a seven year period at the Hastings Reservation, Carmel Valley, California. In: Plumb, Timothy R.; Pillsbury, Norman H., technical coordinators. Proceedings of the symposium on multiple-use management of California's hardwood resources; 1986 November 12-14; San Luis Obispo, CA. Gen. Tech. Rep. PSW-100. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 429-434. [5390]
47. Carmichael, R. S.; Knipe, O. D.; Pase, C. P.; Brady, W. W. 1978. Arizona chaparral: plant associations and ecology. Res. Pap. RM-202. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 16 p. [3038]
48. Chang, Chi-ru. 1996. Ecosystem responses to fire and variations in fire regimes. In: Status of the Sierra Nevada. Sierra Nevada Ecosystem Project: Final report to Congress. Volume II: Assessments and scientific basis for management options. Wildland Resources Center Report No. 37. Davis, CA: University of California, Centers for Water and Wildland Resources: 1071-1099. [28976]
49. Chappell, Christopher B.; Crawford, Rex C.; Barrett, Charley; Kagan, Jimmy; Johnson, David H.; O'Mealy, Mikell; Green, Greg A.; Ferguson, Howard L.; Edge, W. Daniel; Greda, Eva L.; O'Neil, Thomas A. 2001. Wildlife habitats: descriptions, status, trends, and system dynamics. In: Johnson, David H.; O'Neil, Thomas A., managing directors. Wildlife-habitat relationships in Oregon and Washington. Corvallis, OR: Oregon State University Press: 22-114. Available online: http://www.nwhi.org/index/habdescriptions [2008, February 26]. [63870]
50. Chappell, Christopher B.; Giglio, David F. 1999. Pacific madrone forests of the Puget Trough, Washington. In: Adams, A. B.; Hamilton, Clement W., eds. The decline of the Pacific madrone (Arbutus menziesii Pursh): Current theory and research directions: Proceedings of the symposium; 1995 April 28; Seattle, WA. Seattle, WA: Save Magnolia's Madrones, Center for Urban Horticulture, Ecosystems Database Development and Research: 2-11. [40472]
51. Chesnut, V. K. 1902. Plants used by the Indians of Mendocino County, California. Contributions from the U.S. National Herbarium. [Washington, DC]: U.S. Department of Agriculture, Division of Botany. 7(3): 295-408. [54917]
52. Clark, Harold W. 1937. Association types in the North Coast Ranges of California. Ecology. 18: 214-230. [11187]
53. Conard, Susan G. 1987. First year growth of canyon live oak sprouts following thinning and clearcutting. In: Plumb, Timothy R.; Pillsbury, Norman H., technical coordinators. Proceedings of the symposium on multiple-use management of California's hardwood resources; 1986 November 12-14; San Luis Obispo, CA. Gen. Tech. Rep. PSW-100. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 439. [5392]
54. Conard, Susan G.; Emmingham, W. H. 1984. Herbicides for forest brush control in southwestern Oregon. Corvallis, OR: Oregon State University, College of Forestry. 7 p. [10817]
55. Conrad, C. Eugene; Roby, George A.; Hunter, Serena C. 1986. Chaparral and associated ecosystems management: a 5-year research and development program. Gen. Tech. Rep. PSW-91. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 15 p. [4885]
56. Cooper, William Skinner. 1922. The broad-sclerophyll vegetation of California: An ecological study of the chaparral and its related communities. Publ. No. 319. Washington, DC: The Carnegie Institution of Washington. 145 p. [6716]
57. Cornell, Howard V. 1985. Local and regional richness of cynipine gall wasps on California oaks. Ecology. 66(4): 1247-1260. [68675]
58. Dunn, Paul H.; Barro, Susan C.; Wells, Wade G., II; Poth, Mark A.; Wohlgemuth, Peter M.; Colver, Charles G. 1988. The San Dimas Experimental Forest: 50 years of research. Gen. Tech. Rep. PSW-104. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 49 p. [8400]
59. Finch, Sherman J.; McCleery, Dick. 1980. California coast live oak. In: Eyre, F. H., ed. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters: 127-128. [50061]
60. Fites-Kaufmann, Josephine. 1997. Historic landscape pattern and process: fire, vegetation, and environment interactions in the northern Sierra Nevada. Seattle, WA: University of Washington. 175 p. Dissertation. [65695]
61. Flora of North America Association. 2008. Flora of North America: The flora, [Online]. Flora of North America Association (Producer). Available: http://www.fna.org/FNA. [36990]
62. Franklin, Janet; Spears-Lebrun, Linnea A.; Deutschman, Douglas H.; Marsden, Kim. 2006. Impact of a high-intensity fire on mixed evergreen and mixed onifer forests in the Peninsular Ranges of southern California, USA. Forest Ecology and Management. 235(1-3): 18-29. [65016]
63. Franklin, Jerry F.; Dyrness, C. T. 1973. Natural vegetation of Oregon and Washington. Gen. Tech. Rep. PNW-8. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Forest and Range Experiment Station. 417 p. [961]
64. Fried, Jeremy S.; Bolsinger, Charles L.; Beardsley, Debby. 2004. Chaparral in southern and central coastal California in the mid-1990s: area, ownership, condition, and change. Resource Bulletin PNW-RB-240. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 86 p. [50376]
65. Ganey, Joseph L.; Duncan, Russell B.; Block, William M. 1992. Use of oak and associated woodlands by Mexican spotted owls in Arizona. In: Ffolliott, Peter F.; Gottfried, Gerald J.; Bennett, Duane A.; Hernandez C., Victor Manuel; Ortega-Rubio, Alfred; Hamre, R. H., tech. coords. Ecology and management of oak and associated woodlands: perspectives in the southwestern United States and northern Mexico: Proceedings; 1992 April 27-30; Sierra Vista, AZ. Gen. Tech. Rep. RM-218. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 125-128. [22073]
66. Gilbert, Frederick F.; Allwine, Rochelle. 1991. Small mammal communities in the Oregon Cascade Range. In: Ruggiero, Leonard F.; Aubry, Keith B.; Carey, Andrew B.; Huff, Mark H., technical coordinators. Wildlife and vegetation of unmanaged Douglas-fir forests. Gen. Tech. Rep. PNW-GTR-285. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station: 257-267. [17317]
67. Goforth, Brett R.; Graham, Robert C.; Hubbert, Kenneth R.; Zanner, C. William; Minnich, Richard A. 2005. Spatial distribution and properties of ash and thermally altered soils after high-severity forest fire, southern California. International Journal of Wildland Fire. 14: 343-354. [61330]
68. Goudey, Charles B.; Diaz, D.; Chatonian, J.; Frankel, S.; Friday, K.; Husari, S.; Kliejans, J.; Miles, S.; Neisess, J.; Newell, L.; Rose, J. 2004. Section M261E-Sierra Nevada. In: McNab, W. Henry; Avers, Peter E., comps. Ecological subregions of the United States: section descriptions. Administrative Publication WO-WSA-5. Washington, DC: U.S. Department of Agriculture, Forest Service, Ecosystem Management: 33: 5. [64708]
69. Goudey, Charles B.; Diaz, D.; Chatonian, J.; Frankel, S.; Friday, K.; Husari, S.; Kliejans, J.; Miles, S.; Neisess, J.; Newell, L.; Rose, J. 2004. Section M262B-southern California mountains and valleys. In: McNab, W. Henry; Avers, Peter E., comps. Ecological subregions of the United States: section descriptions. Administrative Publication WO-WSA-5. Washington, DC: U.S. Department of Agriculture, Forest Service, Ecosystem Management: 34: 2-3. [64712]
70. Gould, Gordon I., Jr. 1977. Distribution of the spotted owl in California. Western Birds. 8(4): 131-146. [69017]
71. Govaerts, Rafael; Frodin, David G. 1998. World checklist and bibliography of Fagales (Betulaceae, Corylaceae, Fagaceae and Ticodendraceae). Kew, UK: The Royal Botanic Gardens. 497 p. [60947]
72. Graber, David M. 1990. Winter behavior of black bears in the Sierra Nevada, California. In: Darling, Laura M.; Archibald, W. Ralph, eds. Bears--their biology and management: Proceedings, 8th international conference on bear research and management; 1989 February; Victoria, BC. [Place of publication unknown]: International Association for Bear Research and Management: 269-272. [68681]
73. Gratkowski, H. J.; Philbrick, J. R. 1965. Repeated aerial spraying and burning to control sclerophyllous brush. Journal of Forestry. 63(12): 919-923. [8797]
74. Gratkowski, H. 1961. Brush problems in southwestern Oregon. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Forest and Range Experiment Station. 53 p. [8596]
75. Gratkowski, H. 1974. Brushfield reclamation and type conversion. In: Cramer, Owen P., ed. Environmental effects of forest residues management in the Pacific Northwest: A state-of-knowledge compendium. Gen. Tech. Rep. PNW-24. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Forest and Range Experiment Station: I-1 to I-31. [6418]
76. Gratkowski, H. 1978. Herbicides for shrub and weed control in western Oregon. Gen. Tech. Rep. PNW-77. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Forest and Range Experiment Station. 48 p. [6539]
77. Green, Lisle R. 1981. Burning by prescription in chaparral. Gen. Tech. Rep. PSW-51. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 36 p. [19800]
78. Griffin, James R. 1971. Oak regeneration in the upper Carmel Valley, California. Ecology. 52(5): 862-868. [9677]
79. Griffin, James R. 1974. Notes on environment, vegetation and flora: Hastings Natural History Reservation. [Place of publication unknown]: [Publisher unknown]. 90 p. On file at: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory, Missoula, MT. [10531]
80. Griffin, James R. 1978. The Marble-Cone fire ten months later. Fremontia. 6: 8-14. [19081]
81. Griffin, James R.; Critchfield, William B. 1972. The distribution of forest trees in California. Res. Pap. PSW-82. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 118 p. [1041]
82. Gutierrez, R. J.; Koenig, Walter D. 1978. Characteristics of storage trees used by acorn woodpeckers in two California woodlands. Journal of Forestry. 76(3): 162-164. [20555]
83. Gutierrez, R. J.; Verner, Jared; McKelvey, Kevin S.; Noon, Barry R.; Steger, George N.; Call, Douglas R.; LaHaye, William S.; Bingham, Bruce B.; Senser, John S. 1992. Habitat relations of the California spotted owl. In: Verner, Jared; McKelvey, Kevin S.; Noon, Barry R.; Gutierrez, R. J.; Gould, Gordon I., Jr.; Beck, Thomas W., tech. coords. The California spotted owl: a technical assessment of its current status. Gen. Tech. Rep. PSW-GTR-133. Albany, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Research Station: 79-98. [28198]
84. Hann, Wendel; Havlina, Doug; Shlisky, Ayn; [and others]. 2005. Interagency fire regime condition class guidebook. Version 1.2, [Online]. In: Interagency fire regime condition class website. U.S. Department of Agriculture, Forest Service; U.S. Department of the Interior; The Nature Conservancy; Systems for Environmental Management (Producer). Variously paginated [+ appendices]. Available: http://www.frcc.gov/docs/1.2.2.2/Complete_Guidebook_V1.2.pdf [2007, May 23]. [66734]
85. Hanson, Jacob J.; Stuart, John D. 2005. Vegetation responses to natural and salvage logged fire edges in Douglas-fir/hardwood forests. Forest Ecology and Management. 214(1-3): 266-278. [54345]
86. Harrison, Susan. 1997. How natural habitat patchiness affects the distribution of diversity in Californian serpentine chaparral. Ecology. 78(6): 1898-1906. [64476]
87. Hellmers, H.; Horton, J. S.; Juhren, G.; O'Keefe, J. 1955. Root systems of some chaparral plants in southern California. Ecology. 36(4): 667-678. [6147]
88. Hickman, James C., ed. 1993. The Jepson manual: Higher plants of California. Berkeley, CA: University of California Press. 1400 p. [21992]
89. Hobbs, Stephen D.; Wearstler, Kenneth A., Jr. 1985. Effects of cutting sclerophyll brush on sprout development and Douglas- fir growth. Forestry Ecology and Management. 13: 69-81. [9690]
90. Holl, Stephen A.; Bleich, Vernon C.; Torres, Steven G. 2004. Population dynamics of bighorn sheep in the San Gabriel Mountains, California, 1967-2002. Wildlife Society Bulletin. 33(2): 412-426. [62025]
91. Holland, Robert F. 1986. Preliminary descriptions of the terrestrial natural communities of California. Sacramento, CA: California Department of Fish and Game. 156 p. [12756]
92. Holmes, Tyson H. 1990. Botanical trends in northern California oak woodland. Rangelands. 12(1): 3-7. [10939]
93. Horton, Jerome S. 1949. Trees and shrubs for erosion control of southern California mountains. Berkeley, CA: U.S. Department of Agriculture, Forest Service, California Forest and Range Experiment Station; California Department of Natural Resources, Division of Forestry. 72 p. [10689]
94. Horton, Jerome S. 1960. Vegetation types of the San Bernardino Mountains. Tech. Pap. No. 44. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 29 p. [10687]
95. Horton, Jerome S.; Wright, John T. 1944. The wood rat as an ecological factor in southern California watersheds. Ecology. 25(3): 341-351. [10682]
96. Huff, Mark H.; Seavy, Nathaniel E.; Alexander, John D.; Ralph, C. John. 2005. Fire and birds in maritime Pacific Northwest. In: Saab, Victoria A.; Powell, Hugh D. W., eds. Fire and avian ecology in North America. Studies in Avian Biology No. 30. Ephrata, PA: Cooper Ornithological Society: 46-62. [65143]
97. Hunter, J. C.; Parker, V. T.; Barbour, M. G. 1999. Understory light and gap dynamics in an old-growth forested watershed in coastal California. Madrono. 46(1): 1-6. [36548]
98. Hunter, John C. 1997. Correspondence of environmental tolerance with leaf and branch attributes for six co-occurring species of broadleaf evergreen trees in northern California. Trees. 11: 169-175. [27687]
99. Jimerson, Thomas M.; Carothers, Sydney K. 2002. Northwest California oak woodlands: environment, species composition, and ecological status. In: Standiford, Richard B.; McCreary, Douglas; Purcell, Kathryn L., technical coordinators. Proceedings of the 5th symposium on oak woodlands: oaks in California's changing landscape; 2001 October 22-25; San Diego, CA. Gen. Tech. Rep. PSW-GTR-184. Albany, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Research Station: 705-717. [42366]
100. Kartesz, John T. 1999. A synonymized checklist and atlas with biological attributes for the vascular flora of the United States, Canada, and Greenland. 1st ed. In: Kartesz, John T.; Meacham, Christopher A. Synthesis of the North American flora (Windows Version 1.0), [CD-ROM]. Chapel Hill, NC: North Carolina Botanical Garden (Producer). In cooperation with: The Nature Conservancy; U.S. Department of Agriculture, Natural Resources Conservation Service; U.S. Department of the Interior, Fish and Wildlife Service. [36715]
101. Kartesz, John Thomas. 1988. A flora of Nevada. Reno, NV: University of Nevada. 1729 p. [In 2 volumes]. Dissertation. [42426]
102. Kauffman, John Boone. 1986. The ecological response of the shrub component to prescribed burning in mixed conifer ecosystems. Berkeley, CA: University of California. 235 p. Dissertation. [19559]
103. Kearney, Thomas H.; Peebles, Robert H.; Howell, John Thomas; McClintock, Elizabeth. 1960. Arizona flora. 2nd ed. Berkeley, CA: University of California Press. 1085 p. [6563]
104. Keeley, Jon E. 1987. Role of fire in seed germination of woody taxa in California chaparral. Ecology. 68(2): 434-443. [5403]
105. Keeley, Jon E. 2006. South Coast bioregion. In: Sugihara, Neil G.; van Wagtendonk, Jan W.; Shaffer, Kevin E.; Fites-Kaufman, Joann; Thode, Andrea E., eds. Fire in California's ecosystems. Berkeley, CA: University of California Press: 350-390. [65557]
106. Keeley, Jon E.; Zedler, Paul H. 1978. Reproduction of chaparral shrubs after fire: a comparison of sprouting and seeding strategies. The American Midland Naturalist. 99(1): 142-161. [4610]
107. Kennedy, Peter G. 2005. Post-dispersal seed predation varies by habitat not acorn size for Quercus chrysolepis (Fagaceae) and Lithocarpus densiflora (Fagaceae) in central coastal California. Madrono. 52(1): 30-34. [61482]
108. Kilgore, Bruce M. 1971. Response of breeding bird populations to habitat changes in a giant sequoia forest. The American Midland Naturalist. 85(1): 135-152. [7281]
109. Kilgore, Bruce M. 1973. The ecological role of fire in Sierran conifer forests: Its application to National Park management. Quaternary Research. 3: 496-513. [6267]
110. Klyver, F. D. 1931. Major plant communities in a transect of the Sierra Nevada Mountains of California. Ecology. 12(1): 1-17. [+ maps]. [68678]
111. Knops, Johannes M. H.; Koenig, Walter D. 1994. Water use strategies of five sympatric species of Quercus in central coastal California. Madrono. 41(4): 290-301. [40867]
112. Knops, Johannes M. H.; Koenig, Walter D. 1997. Site fertility and leaf nutrients of sympatric evergreen and deciduous species of Quercus in central coastal California. Plant Ecology. 130(2): 121-131. [40862]
113. Koenig, Walter D.; Carmen, William J.; Stanback, Mark T.; Mumme, Ronald L. 1991. Determinants of acorn productivity among five species of oaks in central coastal California. In: Standiford, Richard B., technical coordinator. Proceedings of the symposium on oak woodlands and hardwood rangeland management; 1990 October 31 - November 2; Davis, CA. Gen. Tech. Rep. PSW-126. Berkeley, CA: Pacific Southwest Research Station: 136-142. [18900]
114. Koenig, Walter D.; Heck, M. Katy. 1988. Ability of two species of oak woodland birds to subsist on acorns. The Condor. 90(3): 705-708. [40861]
115. Koenig, Walter D.; Knops, Jean. 1995. Why do oaks produce boom-and-bust seed crops? California Agriculture. 49(5): 7-12. [26010]
116. Koenig, Walter D.; Knops, Johannes M. H. 2002. The behavioral ecology of masting in oaks. In: McShea, William J.; Healy, William M., eds. Oak forest ecosystems: Ecology and management for wildlife. Baltimore, MD: The Johns Hopkins University Press: 129-148. [43526]
117. Koenig, Walter D.; Knops, Johannes M. H.; Carmen, William J.; Stanback, Mark T.; Mumme, Ronald L. 1996. Acorn production by oaks in central coastal California: influence of weather at three levels. Canadian Journal of Forest Research. 26: 1677-1683. [27154]
118. Koenig, Walter D.; Mumme, Ronald L.; Carmen, William J.; Stanback, Mark T. 1994. Acorn production by oaks in central coastal California: variation within and among years. Ecology. 75(1): 99-109. [22738]
119. Koenig, Walter D.; Williams, Pamela L. 1979. Notes on the status of acorn woodpeckers in central Mexico. The Condor. 81(3): 317-318. [68672]
120. Kruckeberg, A. R. 1982. Gardening with native plants of the Pacific Northwest. Seattle, WA: University of Washington Press. 252 p. [9980]
121. Kufeld, Roland C.; Wallmo, O. C.; Feddema, Charles. 1973. Foods of the Rocky Mountain mule deer. Res. Pap. RM-111. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 31 p. [1387]
122. LANDFIRE Rapid Assessment. 2005. Reference condition modeling manual (Version 2.1), [Online]. In: LANDFIRE. Cooperative Agreement 04-CA-11132543-189. Boulder, CO: The Nature Conservancy; U.S. Department of Agriculture, Forest Service; U.S. Department of the Interior (Producers). 72 p. Available: http://www.landfire.gov/downloadfile.php°File=RA_Modeling_Manual_v2_1.pdf [2007, May 24]. [66741]
123. LANDFIRE Rapid Assessment. 2007. Rapid assessment reference condition models, [Online]. In: LANDFIRE. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Lab; U.S. Geological Survey; The Nature Conservancy (Producers). Available: http://www.landfire.gov/models_EW.php [2008, April 18] [66533]
124. Lanner, Ronald M. 1983. Trees of the Great Basin: A natural history. Reno, NV: University of Nevada Press. 215 p. [1401]
125. Lathrop, Earl W.; Martin, Bradford D. 1982. Response of understory vegetation to prescribed burning in yellow pine forests of Cuyamaca Rancho State Park, California. Aliso. 10(2): 329-343. [15943]
126. Lawrence, George; Biswell, Harold. 1972. Effect of forest manipulation on deer habitat in giant sequoia. Journal of Wildlife Management. 36(2): 595-605. [41671]
127. Leach, Howard R. 1956. Food habits of the Great Basin deer herds of California. California Fish and Game. 38: 243-308. [3502]
128. Lininger, Jay C. 2004. Fire history and need for fuel management in mixed Douglas-fir forests of the Klamath-Siskiyou Region, northwest California and southwest Oregon, USA. In: 2nd international wildland fire ecology and fire management congress; 5th symposium on fire and forest meteorology: Proceedings; 2003 November 16-20; Orlando, FL. [Boston, MA]: [American Meteorological Society]: 1-18. [64193]
129. Little, Elbert L., Jr. 1979. Checklist of United States trees (native and naturalized). Agric. Handb. 541. Washington, DC: U.S. Department of Agriculture, Forest Service. 375 p. [2952]
130. Litton, R. Burton. 1980. Oaks and the California landscape. In: Plumb, Timothy R., technical coordinator. Proceedings of the symposium on the ecology, management, and utilization of California oaks; 1979 June 26-28; Claremont, CA. Gen. Tech. Rep. PSW-44. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 161-166. [7031]
131. Lotan, James E.; Alexander, Martin E.; Arno, Stephen F.; French, Richard E.; Langdon, O. Gordon; Loomis, Robert M.; Norum, Rodney A.; Rothermel, Richard C.; Schmidt, Wyman C.; van Wagtendonk, Jan. 1981. Effects of fire on flora: A state-of-knowledge review: National fire effects workshop: Proceedings. 1978 April 10-14; Denver, CO. Gen. Tech. Rep. WO-16. Washington, DC: U.S. Department of Agriculture, Forest Service. 71 p. [1475]
132. Mackie, W. W. 1903. The value of oak leaves for forage. Bulletin No. 150. Berkeley, CA: University of California, College of Agriculture, Agricultural Experiment Station. 21 p. [52010]
133. Mallory, James I. 1980. Canyon live oak. In: Eyre, F. H., ed. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters: 125-126. [7608]
134. Marcot, Bruce G. 1983. Snag use by birds in Douglas-fir clearcuts. In: Davis, Jerry W.; Goodwin, Gregory A.; Ockenfeis, Richard A., technical coordinators. Snag habitat management: proceedings of the symposium; 1983 June 7-9; Flagstaff, AZ. Gen. Tech. Rep. RM-99. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 134-139. [17828]
135. Martin, Bradford D. 1981. Vegetation responses to prescribed burning in a mixed-conifer woodland, Cuyamaca Rancho State Park, California. Loma Linda, CA: Loma Linda University. 112 p. Thesis. [64684]
136. Matsuda, Kozue; McBride, Joe R. 1987. Germination and shoot development of seven California oaks planted at different elevations. In: Plumb, Timothy R.; Pillsbury, Norman H., technical coordinators. Proceedings of the symposium on multiple-use management of California's hardwood resources; 1986 November 12-14; San Luis Obispo, CA. Gen. Tech. Rep. PSW-100. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 79-85. [5400]
137. Matsuda, Kozue; McBride, Joe R. 1989. Germination characteristics of selected California oak species. The American Midland Naturalist. 122: 66-76. [8052]
138. McClaran, Mitchel P. 1987. Yearly variation of blue oak seedling emergence in northern California. In: Plumb, Timothy R.; Pilsbury, Norman H., technical coordinators. Proceedings of the symposium on multiple-use management of California's hardwood resources; 1986 November 12-14; San Luis Obispo, CA. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 76-78. [5531]
139. McDonald, Philip M. 1990. Pseudotsuga macrocarpa (Vasey) Mayr bigcone Douglas-fir. In: Burns, Russell M.; Honkala, Barbara H., technical coordinators. Silvics of North America. Volume 1. Conifers. Agric. Handb. 654. Washington, DC: U.S. Department of Agriculture, Forest Service: 520-526. [13412]
140. McDonald, Philip M.; Fiddler, Gary O. 2003. Effect of chemical and manual release on live oak sprouts and other vegetation in a ponderosa pine plantation on a poor site. Western Journal of Applied Forestry. 18(1): 22-28. [45084]
141. McDonald, Philip M.; Littrell, Edward E. 1976. The bigcone Douglas-fir--canyon live oak community in southern California. Madrono. 23(6): 310-320. [10662]
142. McDonald, Philip M.; Minore, Don; Atzet, Tom. 1983. Southwestern Oregon-northern California hardwoods. In: Burns, Russell M., comp. Silvicultural systems for the major forest types of the United States. Agric. Handb. 445. Washington, DC: U.S. Department of Agriculture: 29-32. [7142]
143. Miller, Howard A.; Lamb, Samuel H. 1985. Oaks Of North America. Happy Camp, CA: Naturegraph Publishers, Inc. 327 p. [68677]
144. Miller, Melanie. 2000. Fire autecology. In: Brown, James K.; Smith, Jane Kapler, eds. Wildland fire in ecosystems: Effects of fire on flora. Gen. Tech. Rep. RMRS-GTR-42-vol. 2. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 9-34. [36981]
145. Minnich, R. A.; Barbour, M. G.; Burk, J. H.; Sosa-Ramirez, J. 2000. California mixed-conifer forests under unmanaged fire regimes in the Sierra San Pedro Martir, Baja California, Mexico. Journal of Biogeography. 27(1): 105-129. [38479]
146. Minnich, R.; Howard, L. 1984. Biogeography and prehistory of shrublands. In: DeVries, Johannes J., ed. Shrublands in California: literature review and research needed for management. Contribution No. 191. Davis, CA: University of California, Water Resources Center: 8-24. [4998]
147. Minnich, Richard A. 1977. The geography of fire and big-cone Douglas-fir, Coulter pine and western conifer forests in the east Transverse Ranges, southern California. In: Mooney, Harold A.; Conrad, C. Eugene, technical coordinators. Proceedings of the symposium on the environmental consequences of fire and fuel management in Mediterranean ecosystems; 1977 August 1-5; Palo Alto, CA. Gen. Tech. Rep. WO-3. Washington, DC: U.S. Department of Agriculture, Forest Service: 443-450. [4875]
148. Minnich, Richard A. 1980. Wildfire and the geographic relationships between canyon live oak, Coulter pine, and bigcone Douglas-fir forests. In: Plumb, Timothy R., technical coordinator. Proceedings of the symposium on the ecology, management and utilization of California oaks; 1979 June 26-28; Claremont, CA. Gen. Tech. Rep. PNW-44. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 55-61. [7015]
149. Minnich, Richard A. 1987. Fire behavior in southern California chaparral before fire control: the Mount Wilson burns at the turn of the century. Annals of the Association of American Geographers. 77(4): 599-618. [28931]
150. Minnich, Richard A. 1987. The distribution of forest trees in northern Baja California, Mexico. Madrono. 34(2): 98-127. [6985]
151. Minnich, Richard A. 1988. The biogeography of fire in the San Bernardino Mountains of California: A historical study. University of California publications in geography: Volume 28. Berkeley, CA: University of California Press. 161 p. [69040]
152. Minnich, Richard A. 1999. Vegetation, fire regimes, and forest dynamics. In: Miller, P. R.; McBride, J. R., eds. Oxidant air pollution impacts in the montane forests of southern California: a case study of the San Bernardino Mountains. Ecological studies: Analysis and synthesis, Vol. 134. New York: Springer-Verlag: 44-80. [30370]
153. Minnich, Richard A.; Everett, Richard G. 2001. Conifer tree distributions in southern California. Madrono. 48(3): 177-197. [40736]
154. Minnich, Richard A.; Franco-Vizcaino, Ernesto. 1997. Mediterranean vegetation of northern Baja California. Fremontia. 25(3): 3-12. [40196]
155. Mirov, N. T.; Kraebel, C. J. 1937. Collecting and propagating the seeds of California wild plants. Res. Note No. 18. Berkeley, CA: U.S. Department of Agriculture, Forest Service, California Forest and Range Experiment Station. 27 p. [9787]
156. Morrison, Michael L.; Block, William M.; Verner, Jared. 1991. Wildlife-habitat relationships in California's oak woodlands: Where do we go from here? In: Standiford, Richard B., technical coordinator. Proceedings of the symposium on oak woodlands and hardwood rangeland management; 1990 October 31 - November 2; Davis, CA. Gen. Tech. Rep. PSW-126. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Research Station: 105-109. [18897]
157. Myatt, Rodney G. 1980. Canyon live oak vegetation in the Sierra Nevada. In: Plumb, Timothy R., technical coordinator. Proceedings of the symposium on the ecology, management and utilization of California oaks; 1979 June 26-28; Claremont, CA. Gen. Tech. Rep. PSW-44. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 86-91. [7019]
158. Nagel, Thomas A.; Taylor, Alan H. 2005. Fire and persistence of montane chaparral in mixed conifer forest landscapes in the northern Sierra Nevada, Lake Tahoe Basin, California, USA. Journal of the Torrey Botanical Society. 132(3): 442-457. [61027]
159. Narog, Marcia G.; Paysen, Timothy E.; Corcoran, Bonni M.; Zavala, Miguel A. 1997. Monitoring fire injury in canyon live oak with electrical resistance. In: Pillsbury, Norman H.; Verner, Jared; Tietje, William D., technical coordinators. Proceedings of a symposium on oak woodlands: ecology, management, and urban interface issues; 1996 March 19-22; San Luis Obispo, CA. Gen. Tech. Rep. PSW-GTR-160. Albany, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Research Station: 646-647. [29047]
160. Narog, Marcia G.; Wilson, Ruth C.; Paysen, Timothy E. 1993. Invertebrate responses to thinning and understory burning in a canyon live oak forest. In: ICFVM, Proceedings of the international conference on forest vegetation management. Auburn, AL: Auburn University School of Forestry Report No. 1: 209-213. [68690]
161. Narog, Maria G.; Paysen, Timothy E. 1994. Multiple-use management of canyon live oak forests using thinning and prescribed burning. Proceedings, 15th annual forest vegetation management conference; 1994 January 25-27; Redding, CA. Redding, CA: Forest Vegetation Management Conference: 154-160. [68696]
162. Neal, Donald L. 1980. Blue oak-digger pine. In: Eyre, F. H., ed. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters: 126-127. [50060]
163. Norman, Steve; Taylor, Alan H. 2002. Variation in fire-return intervals across a mixed-conifer forest landscape. In: Sugihara, Neil G.; Morales, Maria; Morales, Tony, eds. Fire in California ecosystems: integrating ecology, prevention, and management: Proceedings of the symposium; 1997 November 17-20; San Diego, CA. Misc. Publ. No. 1. Davis, CA: Association for Fire Ecology: 170-179. [46056]
164. North, Malcolm; Chen, Jiquan; Oakley, Brian; Song, Bo; Rudnicki, Mark; Gray, Andrew; Innes, Jim. 2004. Forest stand structure and pattern of old-growth western hemlock/Douglas-fir and mixed-conifer forests. Forest Science. 50(3): 299-311. [27077]
165. O'Malley, Penelope Grenoble. 1991. Large-scale restoration on Santa Catalina Island, California. Restoration and Management Notes. 9(1): 7-15. [15763]
166. Parsons, David J. 1981. The historical role of fire in the foothill communities of Sequoia National Park. Madrono. 28(3): 111-120. [13586]
167. Parsons, David J.; DeBenedetti, Steven H. 1979. Impact of fire suppression an a mixed-conifer forest. Forest Ecology and Management. 2: 21-33. [7618]
168. Pavlik, Bruce M.; Muick, Pamela C.; Johnson, Sharon G.; Popper, Marjorie. 1991. Oaks of California. Los Olivos, CA: Cachuma Press, Inc. 184 p. [21059]
169. Paysen, Timothy E.; Ansley, R. James; Brown, James K.; Gottfried, Gerald J.; Haase, Sally M.; Harrington, Michael G.; Narog, Marcia G.; Sackett, Stephen S.; Wilson, Ruth C. 2000. Fire in western shrubland, woodland, and grassland ecosystems. In: Brown, James K.; Smith, Jane Kapler, eds. Wildland fire in ecosystems: Effects of fire on flora. Gen. Tech. Rep. RMRS-GTR-42-vol. 2. Ogden, UT: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station: 121-159. [36978]
170. Paysen, Timothy E.; Narog, Marcia G. 1990. Selective mortality with prescribed fire in canyon live oak. In: Krammes, J. S., technical coordinator. Effects of fire management of southwestern natural resources: Proceedings of the symposium; 1988 November 15-17; Tucson, AZ. Gen. Tech. Rep. RM-191. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 241-243. [11297]
171. Paysen, Timothy E.; Narog, Marcia G. 1993. Tree mortality 6 years after burning a thinned Quercus chrysolepis stand. Canadian Journal of Forest Research. 23(10): 2236-2241. [22734]
172. Paysen, Timothy E.; Narog, Marcia G.; Tissell, Robert G.; Lardner, Melody A. 1991. Trunk and root sprouting on residual trees after thinning a Quercus chrysolepis stand. Forest Science. 37(1): 17-27. [15040]
173. Peinado, M.; Aguirre, J. L.; Delgadillo, J. 1997. Phytosociological, bioclimatic and biogeographical classification of woody climax communities of western North America. Journal of Vegetation Science. 8: 505-528. [28564]
174. Pillsbury, Norman H.; Bonner, Lawrence E.; Thompson, Richard P.; Mark, Walter R.; Cuzick, Roy D. 2004. Long-term growth, sudden oak death assessment and economic viability of coast live oak in three California counties: Seventeen-year results. Volume 1. Technical Report No. 12. San Luis Obispo, CA: California Polytechnic State University, Natural Resources Management Department, Urban Forest Ecosystems Institute. 54 p. [1599]
175. Plumb, T. R. 1961. Sprouting of chaparral by December after a wildfire in July. Technical Paper 57. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 12 p. [9799]
176. Plumb, Tim R. 1980. Response of oaks to fire. In: Plumb, Timothy R., technical coordinator. Proceedings of the symposium on the ecology, management, and utilization of California oaks; 1979 June 26-28; Claremont, CA. Gen. Tech. Rep. PSW-44. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 202-215. [7039]
177. Plumb, Tim R.; Hannah, Bennie. 1991. Artificial regeneration of blue and coast live oaks in the Central Coast. In: Standiford, Richard B., technical coordinator. Proceedings of the symposium on oak woodlands and hardwood rangeland management; 1990 October 31 - November 2; Davis, CA. Gen. Tech. Rep. PSW-126. Berkeley, CA: Pacific Southwest Research Station: 74-80. [18893]
178. Plumb, Timothy R. 1982. Factors affecting germination of southern California oaks. In: Conrad, C. Eugene; Oechel, Walter C., technical coordinators. Proceedings of the symposium on dynamics and management of Mediterranean-type ecosystems; 1981 June 22-26; San Diego, CA. Gen. Tech. Rep. PSW-58. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 625. [6093]
179. Plumb, Timothy R.; Gomez, Anthony P. 1983. Five southern California oaks: identification and postfire management. Gen. Tech. Rep. PSW-71. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 56 p. [5898]
180. Plumb, Timothy R.; McDonald, Philip M. 1981. Oak management in California. Gen. Tech. Rep. PSW-54. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 11 p. [6568]
181. Raabe, Robert D. 1980. Diseases of oaks in California. In: Plumb, Timothy R., tech. coord. Proceedings of the symposium on the ecology, management, and utilization of California oaks; 1979 June 26-28; Claremont, CA. Gen. Tech. Rep. PSW-44. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 195-201. [7038]
182. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843]
183. Regelbrugge, Jon C.; Conard, Susan G. 1993. Modeling tree mortality following wildfire in Pinus ponderosa forests in the central Sierra Nevada of California. International Journal of Wildland Fire. 3(3): 139-148. [22044]
184. Riggan, Philip J.; Dunn, Paul H. 1982. Harvesting chaparral biomass for energy--an environmental assessment. In: Conrad, C. Eugene; Oechel, Walter C., technical coordinators. Proceedings of the symposium on dynamics and management of Mediterranean-type ecosystems; 1981 June 22-26; San Diego, CA. Gen. Tech. Rep. PSW-58. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 149-157. [6019]
185. Rizzo, David M.; Garbelotto, Matteo. 2003. Sudden oak death: endangering California and Oregon forest ecosystems. Frontiers in Ecology and the Environment. 1(4): 197-204. [68680]
186. Robinson, Cyril S. 1937. Plants eaten by California mule deer on the Los Padres National Forest. Journal of Forestry. 35(3): 285-292. [51853]
187. Rossi, Randall S. 1980. History of cultural influences on the distribution and reproduction of oaks in California. In: Plumb, Timothy R., technical coordinator. Proceedings of the symposium on the ecology, management and utilization of California oaks; 1979 June 26-28; Claremont, CA. Gen. Tech. Rep. PSW-44. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 7-18. [7010]
188. Rundel, Philip W. 1987. Origins and adaptations of California hardwoods. In: Plumb, Timothy R.; Pillsbury, Norman H., technical coordinators. Proceedings of the symposium on multiple-use management of California's hardwood resources; 1986 November 12-14; San Luis Obispo, CA. Gen. Tech. Rep. PSW-100. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station: 11-17. [5355]
189. Sampson, Arthur W.; Jespersen, Beryl S. 1963. California range brushlands and browse plants. Berkeley, CA: University of California, Division of Agricultural Sciences, California Agricultural Experiment Station, Extension Service. 162 p. [3240]
190. Savage, Melissa. 1994. Anthropogenic and natural disturbance and patterns of mortality in a mixed conifer forest in California. Canadian Journal of Forest Research. 24(6): 1149-1159. [64659]
191. Sawyer, John O., Jr. 1980. Douglas-fir-tanoak-Pacific madrone. In: Eyre, F. H., ed. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters: 111-112. [50048]
192. Sawyer, John O., Jr.; Keeler-Wolf, Todd. 1997. A manual of California vegetation, [Online]. [Sacramento, CA]: California Native Plant Society (Producer). Available: http://davisherb.ucdavis.edu/cnpsActiveServer/index.html [2008, January 31]. [48720]
193. Sawyer, John O.; Sillett, Stephen C.; Popenoe, James H.; LaBanca, Anthony; Sholars, Teresa; Largent, David L.; Euphrat, Fred; Noss, Reed F.; Van Pelt, Robert. 2000. Characteristics of redwood forests. In: Noss, Reed F., ed. The redwood forest: History, ecology, and conservation of the coast redwoods. Washington, DC: Island Press: 39-79. [40464]
194. Sawyer, John O.; Thornburgh, Dale A.; Griffin, James R. 1977. Mixed evergreen forest. In: Barbour, Michael G.; Major, Jack, eds. Terrestrial vegetation of California. New York: John Wiley and Sons: 359-381. [7218]
195. Schimke, Harry E.; Green, Lisle R. 1970. Prescribed fire for maintaining fuel-breaks in the central Sierra Nevada. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 9 p. [11189]
196. Shreve, Forrest. 1927. The vegetation of a coastal mountain range. Ecology. 8(1): 27-44. [63417]
197. Skinner, Carl N.; Chang, Chi-ru. 1996. Fire regimes, past and present. In: Status of the Sierra Nevada. Sierra Nevada Ecosystem Project: Final report to Congress. Volume II: Assessments and scientific basis for management options. Wildland Resources Center Report No. 37. Davis, CA: University of California, Centers for Water and Wildland Resources: 1041-1069. [28975]
198. Skinner, Carl N.; Taylor, Alan H. 2006. Southern Cascades bioregion. In: Sugihara, Neil G.; van Wagtendonk, Jan W.; Shaffer, Kevin E.; Fites-Kaufman, Joann; Thode, Andrea E., eds. Fire in California's ecosystems. Berkeley, CA: University of California Press: 195-224. [65540]
199. Skinner, Carl N.; Taylor, Alan H.; Agee, James K. 2006. Klamath Mountains bioregion. In: Sugihara, Neil G.; van Wagtendonk, Jan W.; Shaffer, Kevin E.; Fites-Kaufman, Joann; Thode, Andrea E., eds. Fire in California's ecosystems. Berkeley, CA: University of California Press: 170-194. [65539]
200. Standiford, Richard; McDougald, Neil; Frost, William; Phillips, Ralph. 1997. Factors influencing the probability of oak regeneration on southern Sierra Nevada woodlands in California. Madrono. 44(2): 170-183. [51873]
201. Steger, George N.; Eberlein, Gary E.; Munton, Thomas E.; Johnson, Kenneth D. 1997. Characteristics of California spotted owl nests in foothill riparian and oak woodlands of the southern Sierra Nevada, California. In: Pillsbury, Norman H.; Verner, Jared; Tietje, William D., technical coordinators. Proceedings of a symposium on oak woodlands: ecology, management, and urban interface issues; 1996 March 19-22; San Luis Obispo, CA. Gen. Tech. Rep. PSW-GTR-160. Albany, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Research Station: 355-364. [29027]
202. Stein, William I. 1980. Oregon white oak. In: Eyre, F. H., ed. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters: 110-111. [9857]
203. Stephenson, John R.; Calcarone, Gena M. 1999. Mountain and foothills ecosystems: habitat and species conservation issues. In: Stephenson, John R.; Calcarone, Gena M. Southern California mountains and foothills assessment. Gen. Tech. Rep. PSW-GTR-172. Albany, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Research Station: 15-60. [35514]
204. Stewart, Glenn R.; Jennings, Mark R.; Goodman, Robert H., Jr. 2005. Sensitive species of snakes, frogs, and salamanders in southern California conifer forest areas: status and management. In: Kus, Barbara E.; Beyers, Jan L., tech. coords. Planning for biodiversity: Bringing research and management together: Proceedings of a symposium for the south coast ecoregion; 2000 February 29 - March 2; Pomona, CA. Gen. Tech. Rep. PSW-GTR-195. Albany, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Research Station: 165-197. [64510]
205. Stickney, Peter F. 1989. Seral origin of species comprising secondary plant succession in Northern Rocky Mountain forests. FEIS workshop: Postfire regeneration. Unpublished draft on file at: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory, Missoula, MT. 10 p. [20090]
206. Stratton, Lisa. 2002. Oak restoration trials: Santa Catalina Island. In: Standiford, Richard B.; McCreary, Douglas; Purcell, Kathryn L., technical coordinators. Proceedings of the 5th symposium on oak woodlands: oaks in California's changing landscape; 2001 October 22-25; San Diego, CA. Gen. Tech. Rep. PSW-GTR-184. Albany, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Research Station: 837-838. [42381]
207. Strothmann, R. O.; Roy, Douglass F. 1984. Regeneration of Douglas-fir in the Klamath Mountains Region, California and Oregon. Gen. Tech. Rep. PSW-81. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 35 p. [5640]
208. Stuart, John D.; Grifantini, Michael C.; Fox, Lawrence, III. 1993. Early successional pathways following wildfire and subsequent silvicultural treatment in Douglas-fir/hardwood forests, nw California. Forest Science. 39(3): 561-572. [22064]
209. Stuart, John D.; Stephens, Scott L. 2006. North Coast bioregion. In: Sugihara, Neil G.; van Wagtendonk, Jan W.; Shaffer, Kevin E.; Fites-Kaufman, Joann; Thode, Andrea E., eds. Fire in California's ecosystems. Berkeley, CA: University of California Press: 147-169. [65538]
210. Stubblefield, Cynthia H. 1993. Food habits of black bear in the San Gabriel Mountains of southern California. The Southwestern Naturalist. 38(3): 290-293. [22146]
211. Swetnam, Thomas W.; Touchan, Ramzi; Baisan, Christopher H.; Caprio, Anthony C.; Browns, Peter M. 1991. Giant sequoia fire history in Mariposa Grove, Yosemite National Park. In: Yosemite centennial symposium proceedings: natural areas and Yosemite: prospects for the future: a global issues symposium joining the 17th Annual Natural Areas Conference with the Yosemite Centennial Celebration; 1990 October 13-20; Denver, CO. NPS D-374. Denver, CO: National Park Service, Branch of Publications and Graphic Design, Denver Service Center: 249-255. [52904]
212. Taber, Richard D.; Dasmann, Raymond F. 1958. The black-tailed deer of the chaparral: Its life history and management in the north Coast Range of California. Game Bulletin No. 8. Sacramento, CA: State of California, Department of Fish and Game, Game Management Branch. 166 p. [16312]
213. Talley, Steven N.; Griffin, James R. 1980. Fire ecology of a montane pine forest, Junipero Serra Peak, California. Madrono. 27: 49-60. [4788]
214. Talley, Steven Neal. 1974. The ecology of Santa Lucia fir (Abies bracteata), a narrow endemic of California. Durham, NC: Duke University. 208 p. Dissertation. [23160]
215. Tappeiner, John C., II; McDonald, Philip M.; Newton, Michael; Harrington, Timothy B. 1992. Ecology of hardwoods, shrubs, and herbaceous vegetation: effects on conifer regeneration. In: Hobbs, Stephen D.; Tesch, Steven D.; Owston, Peyton W.; Stewart, Ronald E.; Tappeiner, John C., II; Wells, Gail E., eds. Reforestation practices in southwestern Oregon and northern California. Corvallis, OR: Oregon State University, Forest Research Laboratory: 136-164. [22157]
216. Taylor, Alan H.; Skinner, Carl N. 2003. Spatial patterns and controls on historical fire regimes and forest structure in the Klamath Mountains. Ecological Applications. 13(3): 704-719. [52969]
217. Tesch, Steven D.; Hobbs, Stephen D. 1986. Sprouting brush is tough competition for planted Douglas-fir seedlings in southwest Oregon. In: Proceedings, 7th annual forest vegetation management conference; 1985 November 6-7; Eureka, CA. Redding, CA: Forest Vegetation Management Conference: 81-84. [3839]
218. Tesch, Steven D.; Hobbs, Stephen D. 1989. Impact of shrub sprout competition on Douglas-fir seedling development. Western Journal of Applied Forestry. 4(3): 89-92. [7615]
219. Thornburgh, Dale A. 1990. Quercus chrysolepis Liebm. canyon live oak. In: Burns, Russell M.; Honkala, Barbara H., tech. coords. Silvics of North America. Vol. 2. Hardwoods. Agric. Handbook 654. Washington, DC: U.S. Department of Agriculture, Forest Service: 618-624. [10668]
220. Thorne, Robert F. 1977. Montane and subalpine forests of the Transverse and Peninsular ranges. In: Barbour, Michael G.; Major, Jack, eds. Terrestrial vegetation of California. New York: John Wiley and Sons: 537-557. [7214]
221. Tucker, John M. 1983. California's native oaks. Fremontia. 11(3): 3-12. [52714]
222. U.S. Department of Agriculture, Forest Service. 1964. California mixed evergreen forest (Quercus-Arbutus-Pseudotsuga). In: Kuchler, A. W. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society: 29. [67226]
223. U.S. Department of Agriculture, Natural Resources Conservation Service. 2008. PLANTS Database, [Online]. Available: https://plants.usda.gov /. [34262]
224. Van Dersal, William R. 1940. Utilization of oaks by birds and mammals. Journal of Wildlife Management. 4(4): 404-428. [11983]
225. van Wagtendonk, J. W. 1991. Spatial analysis of lightning strikes in Yosemite National Park. In: Andrews, Patricia L.; Potts, Donald F., eds. Proceedings, 11th conference on fire and forest meteorology; 1991 April 16-19; Missoula, MT. SAF Publication 91-04. Bethesda, MD: Society of American Foresters: 605-611. [48238]
226. van Wagtendonk, Jan W.; Fites-Kaufman, Joann. 2006. Sierra Nevada bioregion. In: Sugihara, Neil G.; van Wagtendonk, Jan W.; Shaffer, Kevin E.; Fites-Kaufman, Joann; Thode, Andrea E., eds. Fire in California's ecosystems. Berkeley, CA: University of California Press: 264-294. [65544]
227. Vankat, John L.; Major, Jack. 1978. Vegetation changes in Sequoia National Park, California. Journal of Biogeography. 5: 377-402. [17353]
228. Vasek, Frank C.; Clovis, Jesse F. 1976. Growth forms in Arctostaphylos glauca. American Journal of Botany. 63(2): 189-195. [11994]
229. Verner, Jared; McKelvey, Kevin S.; Noon, Barry R; Gutierrez, R. J.; Gould, Gordon I., Jr.; Beck, Thomas W. 1992. Assessment of the current status of the California spotted owl, with recommendations for management. In: Verner, Jared; McKelvey, Kevin S.; Noon, Barry R.; Gutierrez, R. J.; Gould, Gordon I., Jr.; Beck, Thomas W., tech. coords. The California spotted owl: a technical assessment of its current status. Gen. Tech. Rep. PSW-GTR-133. Albany, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Research Station: 3-26. [28195]
230. Vogl, Richard J. 1973. Ecology of knobcone pine in the Santa Ana Mountains, California. Ecological Monographs. 43: 125-143. [4815]
231. Vogl, Richard J. 1976. An introduction to the plant communities of the Santa Ana and San Jacinto Mountains. In: Latting, June, ed. Symposium proceedings: plant communities of southern California; 1974 May 4; Fullerton, CA. Special Publication No. 2. Berkeley, CA: California Native Plant Society: 77-98. [4230]
232. Vogl, Richard J.; Miller, Brian C. 1968. The vegetational composition of the south slope of Mt. Pinos, California. Madrono. 19(7): 225-288. [31340]
233. Wangler, Michael J.; Minnich, Richard A. 1996. Fire and succession in pinyon-juniper woodlands of the San Bernardino Mountains, California. Madrono. 43(4): 493-514. [27891]
234. Waring, R. H. 1969. Forest plants of the eastern Siskiyous: their environment and vegetational distribution. Northwest Science. 43(1): 1-17. [9047]
235. Warner, Thomas E. 1980. Fire history in the yellow pine forest of Kings Canyon National Park. In: Stokes, Marvin A.; Dieterich, John H., technical coordinators. Proceedings of the fire history workshop; 1980 October 20-24; Tucson, AZ. Gen. Tech. Rep. RM-81. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 89-92. [16047]
236. Washington State Cooperative Extension Service. 1982. Herbicides in forestry. Pullman, WA: Washington State University, College of Agriculture, Cooperative Extension Service. 13 p. [7873]
237. Weatherspoon, C. Philip. 1990. Sequoiadendron giganteum (Lindl.) Buchholz. In: Burns, Russell M.; Honkala, Barbara H., technical coordinators. Silvics of North America. Volume 1. Conifers. Agric. Handb. 654. Washington, DC: U.S. Department of Agriculture, Forest Service: 552-562. [13415]
238. Weatherspoon, C. Phillip; Husari, Susan J.; van Wagtendonk, Jan W. 1992. Fire and fuels management in relation to owl habitat in forests of the Sierra Nevada and southern California. In: Verner, Jared; McKelvey, Kevin S.; Noon, Barry R.; Gutierrez, R. J.; Gould, Gordon I., Jr.; Beck, Thomas W., tech. coords. The California spotted owl: a technical assessment of its current status. Gen. Tech. Rep. PSW-GTR-133. Albany, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Research Station: 247-260. [22630]
239. Weise, David R.; Sackett, Stephen S.; Paysen, Timothy E.; Haase, Sally M.; Narog, Marcia G. 1996. Rx fire research for southwestern forests. Fire Management Notes. 56(2): 23-25. [30503]
240. Wells, Philip V. 1962. Vegetation in relation to geological substratum and fire in the San Luis Obispo quadrangle, California. Ecological Monographs. 32(1): 79-103. [14183]
241. Wheeler, Gary P.; Fancher, Jack M. 1984. San Diego County riparian systems: current threats and statutory protection efforts. In: Warner, Richard E.; Hendrix, Kathleen M., eds. California riparian systems: Ecology, conservation, and productive management. Berkeley, CA: University of California Press: 838-843. [5875]
242. White, Diane E.; Atzet, T.; Martinez, P. A. 2003. Vegetation recovery in the Biscuit Fire, Siskiyou National Forest, Oregon. In: 2nd international wildland fire ecology and fire management congress; 5th symposium on fire and forest meteorology: Proceedings; 2003 November 17-20; Orlando, FL. Boston, MA: American Meteorological Society: 76. [Abstract]. Available online: http://ams.confex.com/ams/FIRE2003/techprogram/paper_66934.htm [2005, November 14]. [55125]
243. Whittaker, R. H. 1960. Vegetation of the Siskiyou Mountains, Oregon and California. Ecological Monographs. 30(3): 279-338. [6836]
244. Wieslander, A. E. 1964. California oakwoods (Quercus). In: Kuchler, A. W. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society: 30. [67228]
245. Wiggins, Ira L. 1980. Flora of Baja California. Stanford, CA: Stanford University Press. 1025 p. [21993]
246. Wills, Robin D.; Stuart, John D. 1994. Fire history and stand development of a Douglas-fir/hardwood forest in northern California. Northwest Science. 68(3): 205-211. [23901]
247. Wirtz, William O., II. 1979. Effects of fire on birds in chaparral. In: Koch, David L.; Armstrong, R.; Baker, S.; Lider, E.; Robertson, S.; Vigg, S., eds. California-Nevada Wildlife Transactions; 1979 February 1-3; Long Beach, CA. [Bethesda, MD]: Wildlife Society; American Fisheries Society: 114-124. [54810]
248. Wohlgemuth, Peter M.; Hubbert, Ken R.; Robichaud, Peter R. 2001. The effects of log erosion barriers on post-fire hydrologic response and sediment yield in small forested watersheds, southern California. Hydrological Processes. 15(15): 3053-3066. [64669]
249. Wolf, Carl B. 1945. California wild tree crops. Anaheim, CA: Rancho Santa Ana Botanic Garden: 67 p. [63167]
250. Wright, Henry A.; Bailey, Arthur W. 1982. Fire ecology: United States and southern Canada. New York: John Wiley & Sons. 501 p. [2620]
251. Young, James A.; Evans, Raymond A.; Kay, Burgess L.; Budy, Jerry D. 1986. California black oak woodlands in the Great Basin. Transactions of the Western Section of the Wildlife Society. 22: 117-120. [65869]
252. Zielinski, William J.; Duncan, Neil P.; Farmer, Emma C.; Truex, Richard L.; Clevenger, Anthony P.; Barrett, Reginald H. 1999. Diet of fishers (Martes pennanti) at the southernmost extent of their range. Journal of Mammalogy. 80(3): 961-971. [55515]
253. Zigmond, Maurice L. 1981. Kawaiisu ethnobotany. Salt Lake City, UT: University of Utah Press. 102 p. [35936]