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Horton, Scott P.; Mannan, R. William. 1988. Effects of prescribed fire on snags and cavity-nesting birds in southeastern Arizona pine forests. Wildlife Society Bulletin. 16: 37-44 [2].
SPECIES INCLUDED IN THE SUMMARY:Small snags in unburned stands showed little evidence of decay and contained no cavities. Most large snags in unburned stands showed some evidence of decay:
Density and condition of Arizona pine snags in unburned stands [2] |
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Decay class | Number of snags/ha* | Description of decay class | ||
Limbs | Sapwood | Bark cover (%) | ||
I | <0.1 | Foliage still on twigs | Sound | 100 |
II | 4.0 | Fine twigs remaining | Sound to rotting | 75-100 |
III | 1.9 | Secondary branches remaining | Intact to rotting | 25-85 |
IV | 3.0 | Branch stubs remaining | Some sloughed | 10-50 |
V | 2.5 | None | Much sloughed | 0-50 |
VI | 0.5 | Secondary branches remaining | Sound to rotting | <10 |
*Average of density in control stands and prefire treatment stands |
Densities of cavity-nesting birds before treatment are shown below with posttreatment data (see Fire Effects on Cavity-nesting Birds). In unburned stands, cavity nesters generally preferred snags >50 cm DBH in decay classes II or III, although nest-tree preferences varied for the 8 most common species:
Nest tree preferences* of 8 cavity-nesting bird species in unburned stands [2] |
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Bird species | Preferred size of large snag (cm DBH) | Preferred decay class |
House wren | no preference | no preference |
Mountain chickadee | >30 | II |
Pygmy nuthatch | >50 | III |
Violet-green swallow | 15 to <30 | III |
Western bluebird | no preference | III |
Woodpecker species** | >50 | II |
*Preferences shown in this table are all statistically significant
(P<0.10), except for mountain chickadee (for which all 5 nests occurred
in the same size and decay class) **Data were combined for 3 species: acorn woodpecker, hairy woodpecker, and northern flicker |
In the 65 years prior to this study, no wildfires in the study area were larger than 2 ha. The authors note that, prior to the 20th century, fires were probably frequent and extensive in southwestern ponderosa pine forests (including Arizona pine), burning every 5 to 12 years [2]. A recent synthesis of fire regime information for southwestern ponderosa pine forests [4] provides this description of the historical fire regime:
Fire regime information on the vegetation community studied in this Research Project Summary. Fire regime characteristics are taken from the LANDFIRE Rapid Assessment Vegetation Model [4]. This vegetation model was developed by local experts using available literature, local data, and expert opinion as documented in the PDF file linked from the Potential Natural Vegetation Group listed below. Cells are blank where information is not available in the Rapid Assessment Vegetation Model. | |||||
Vegetation Community (Potential Natural Vegetation Group) | Fire severity* | Fire regime characteristics | |||
Percent of fires | Mean interval (years) |
Minimum interval (years) |
Maximum interval (years) |
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Southwest mixed conifer (warm, dry with aspen) | Replacement | 7% | 300 | ||
Mixed | 13% | 150 | 80 | 200 | |
Surface or low | 80% | 25 | 2 | 70 | |
*Fire Severities: Replacement=Any fire that causes greater than 75% top removal of a vegetation-fuel type, resulting in general replacement of existing vegetation; may or may not cause a lethal effect on the plants. Mixed=Any fire burning more than 5% of an area that does not qualify as a replacement, surface, or low-severity fire; includes mosaic and other fires that are intermediate in effects. Surface or low=Any fire that causes less than 25% upper layer replacement and/or removal in a vegetation-fuel class but burns 5% or more of the area [1,3]. |
Woody fuel reduction planned for prescribed burns in the study [2] |
|
Woody fuel size class | Planned % reduction in fuel load |
<0.6 cm diameter | 70 |
0.6-7.6 cm diameter | 55 |
>7.6 cm diameter | 25 |
Research Objectives: This study compared 3 characteristics of Arizona pine forests before and after prescribed fire: number and characteristics of Arizona pine snags, availability of snags used by cavity-nesting birds, and breeding populations of cavity-nesting birds.
Fires were broadcast underburns that backed downslope and into the wind. Two stands were burned in mid-November 1984; the third was burned in early May 1985. Burns were conducted within the prescribed weather conditions listed below, and fire behavior generally stayed within the prescribed limits.
Range of burning conditions and fire behavior acceptable for prescribed burns [2] | |
Air temperature | 4 to 24 °C |
Relative Humidity | 25% to 40% |
Wind speed | 1-16 km/hr |
Flame height | 0.3-1.0 m |
Flame length | 0.3-1.2 m |
Rate of spread | 40-120 m/hr |
FIRE EFFECTS ON SNAGS:
Fire effects on Arizona pine snags were measured the first summer after prescribed burning (summer 1985).
Treatments reduced both large and small snags by about 50%. Density of the kind of snags generally
preferred by cavity nesters (>50 cm DBH in decay classes
II or III) were reduced from 1.5 to 1.0 snags/ha. Snags with greater decay were significantly
(P<0.001) more vulnerable to fire damage; 73% of snags in decay class IV burned down
or were reduced more than 50% in height, in comparison with less than 20% of snags in decay classes
II and VI. Snags >30 cm DBH were more likely to burn than smaller snags, probably because larger
snags had more advanced decay. Snags with large amounts of undecayed twigs, branches, and sloughed
bark at the base ignited more easily than those with less fuel or mostly decayed fuel at the base.
Few large snags were created by the burns, but small snags increased 20-fold:
Effects of understory prescribed burns on density of Arizona pine snags [2] |
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Size class | Prefire density (number/ha) | Postfire density (number/ha) | ||
Burned* | Created** | Total after fire | ||
Large (>15 cm DBH) | 11.1 | 5.5 | 0.5 | 6.1 |
Small (<15 cm DBH) | 10.9 | 6.0 | 216.1 | 221.0 |
*Burned down, were reduced >50% in
height, or had most of sapwood burned away **Killed by prescribed burns |
FIRE EFFECTS ON CAVITY-NESTING BIRDS:
Cavity-nesting birds were surveyed the first breeding season after prescribed burning
(May-July 1985). Only 3 species showed significant (P<0.10) differences in
density between burned and unburned sites (which included both control sites and preburn
treatment sites): Mountain chickadee density was greater,
and northern flicker and violet-green swallow densities decreased. The authors attributed the differences to changes in prey
populations, shifts in foraging areas after burning, or both.
Density (number/40 ha) of cavity-nesting birds before and after understory fire [2] |
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Unburned (control) sites |
Burned sites |
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Species | 1984 | 1985 | 1984 (prefire) |
1985 (breeding season after treatments) |
Acorn woodpecker | 1.1 | 1.5 | 4.1 | 2.8 |
Ash-throated flycatcher | P* | |||
Bridled titmouse | P | P | P | P |
Bewick's wren | P | P | P | P |
Brown creeper | 5.8 | 7.3 | 9.8 | 7.9 |
Cordilleran flycatcher (western flycatcher) | 62.4 | 48.0 | 39.2 | 67.0 |
Dusky-capped flycatcher | P | P | P | P |
Eastern bluebird | P | |||
Hairy woodpecker | 0.7 | 0.8 | 0.8 | 1.0 |
House wren | 53.2 | 91.8 | 15.0 | 46.2 |
Mountain chickadee | 7.2 | 7.0 | 5.6 | 12.2** |
Northern flicker | 1.7 | 2.3 | 2.6 | 2.1** |
Pygmy nuthatch | 26.2 | 15.8 | 24.4 | 14.2 |
Violet-green swallow | 31.7 | 34.0 | 37.4 | 17.6** |
Western bluebird | 11.3 | 6.1 | 12.5 | 6.2 |
White-breasted nuthatch | 4.4 | 2.9 | 7.0 | 4.2 |
*P=present at low density. Blank cell indicates not present. **Significantly different from density on unburned sites (P<0.10) |
FIRE MANAGEMENT IMPLICATIONS:
Management objectives for the burns, to improve regeneration and growth of
Arizona pines and reduce woody fuels, were met.
The authors recommend that protection of large snags (by constructing fuel breaks around
the trees) be considered before conducting prescribed underburns in forests where large snags
have been reduced by logging. It is most important to protect snags >50 cm DBH in
decay classes II and III, which can provide nest sites for most
cavity-nesting birds, and snags in decay class I, which may provide nest sites in the future.
Common name | Scientific name |
Trees | |
Arizona pine | Pinus arizonica |
Birds | |
Bridled titmouse | Baeolophus wollweberi (Parus wollweberi)* |
Brown creeper | Certhia americana |
Northern flicker | Colaptes auratus |
Cordilleran flycatcher (western flycatcher) | Empidonax occidentalis (Empidonax difficilis) |
Acorn woodpecker | Melanerpes formicivorus |
Ash-throated flycatcher | Myiarchus cinerascens |
Dusky-capped flycatcher | Myiarchus tuberculifer |
Hairy woodpecker | Picoides villosus |
Mountain chickadee | Poecile gambeli (Parus gambeli) |
Eastern bluebird | Sialia sialis |
Western bluebird | Sialia mexicana |
White-breasted nuthatch | Sitta carolinensis |
Pygmy nuthatch | Sitta pygmaea |
Violet-green swallow | Tachycineta thalassina |
Bewick's wren | Thryomanes bewickii |
House wren | Troglodytes aedon |
*For species that have undergone scientific name changes, names in parentheses are those used in the research paper. |
1. Hann, Wendel; Havlina, Doug; Shlisky, Ayn; [and others]. 2005. Interagency fire regime condition class guidebook. Version 1.2, [Online]. In: Interagency fire regime condition class website. U.S. Department of Agriculture, Forest Service; U.S. Department of the Interior; The Nature Conservancy; Systems for Environmental Management (Producer). Variously paginated [+ appendices]. Available: http://www.frcc.gov/docs/1.2.2.2/Complete_Guidebook_V1.2.pdf [2007, May 23]. [66734]
2. Horton, Scott P.; Mannan, R. William. 1988. Effects of prescribed fire on snags and cavity-nesting birds in southeastern Arizona pine forests. Wildlife Society Bulletin. 16: 37-44. [5549]
3. LANDFIRE Rapid Assessment. 2005. Reference condition modeling manual (Version 2.1), [Online]. In: LANDFIRE. Cooperative Agreement 04-CA-11132543-189. Boulder, CO: The Nature Conservancy; U.S. Department of Agriculture, Forest Service; U.S. Department of the Interior (Producers). 72 p. Available: http://www.landfire.gov/downloadfile.php?file=RA_Modeling_Manual_v2_1.pdf [2007, May 24]. [66741]
4. LANDFIRE Rapid Assessment. 2007. Rapid assessment reference condition models. In: LANDFIRE. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Lab; U.S. Geological Survey; The Nature Conservancy (Producers). Available: http://www.landfire.gov/models_EW.php [66533]