Index of Species Information
SPECIES: Quercus coccinea
Introductory
SPECIES: Quercus coccinea
AUTHORSHIP AND CITATION :
Carey, Jennifer H. 1992. Quercus coccinea. In: Fire Effects Information System, [Online].
U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station,
Fire Sciences Laboratory (Producer). Available:
https://www.fs.usda.gov/database/feis/plants/tree/quecoc/all.html [].
ABBREVIATION :
QUECOC
SYNONYMS :
Quercus richteri Baenitz
SCS PLANT CODE :
QUCO2
COMMON NAMES :
scarlet oak
Spanish oak
TAXONOMY :
The currently accepted scientific name of scarlet oak is Quercus coccinea
Muenchh. (Fagaceae)[24,30]. Scarlet oak has been placed within the the
subgenus Erythrobalanus, or red (black) oak group [20]. A rarely recognized
variety, Quercus coccinea var. tuberculata Sarg., is distinguished by
thickened tuberculate scales of the cup [5].
Scarlet oak hybridizes with the following species [24,30]:
x Q. ilicifolia (bear oak): Q. X robbinsii Trel.
x Q. velutina (black oak): Q. X fontana Laughlin
x Q. palustris (pin oak)
LIFE FORM :
Tree
FEDERAL LEGAL STATUS :
No special status
OTHER STATUS :
In Maine's Official List of Endangered and Threatened Plants, scarlet
oak is listed under the administrative category, Special
Concern-Possibly Extirpated [13].
DISTRIBUTION AND OCCURRENCE
SPECIES: Quercus coccinea
GENERAL DISTRIBUTION :
Scarlet oak is distributed from southwestern Maine west to New York,
Ohio, southern Michigan and Indiana; south to southern Illinois,
southeastern Missouri, and central Mississippi; east to southern Alabama
and southwestern Georgia; and north along the western edge of the
Atlantic Coastal Plain to the Virginia Coast. Scarlet oak is abundant
in the Piedmont and in the Appalachian Mountains [24,30].
ECOSYSTEMS :
FRES10 White - red - jack pine
FRES13 Loblolly - shortleaf pine
FRES14 Oak - pine
FRES15 Oak - hickory
STATES :
AL CT DE GA IL IN KY ME MD MA
MI MS MO NH NJ NY NC OH PA RI
SC TN VA VT WV
BLM PHYSIOGRAPHIC REGIONS :
NO-ENTRY
KUCHLER PLANT ASSOCIATIONS :
K095 Great Lakes pine forest
K100 Oak - hickory forest
K104 Appalachian oak forest
K110 Northeastern oak - pine forest
K111 Oak - hickory - pine forest
SAF COVER TYPES :
14 Northern pin oak
40 Post oak - blackjack oak
43 Bear oak
44 Chestnut oak
45 Pitch pine
51 White pine - chestnut oak
52 White oak - black oak - northern red oak
53 White oak
75 Shortleaf pine
76 Shortleaf pine - oak
78 Virginia pine - oak
79 Virginia pine
82 Loblolly pine - hardwood
110 Black oak
SRM (RANGELAND) COVER TYPES :
NO-ENTRY
HABITAT TYPES AND PLANT COMMUNITIES :
Scarlet oak is a common component of many eastern and central dry upland
forests. Nearly pure stands of scarlet oak grow in areas of the Ozark
Plateau in Missouri [24]. A chestnut oak (Quercus prinus)-scarlet oak
variant of the chestnut oak SAF cover type is found on upper slopes and
ridges in the central Appalachians. Scarlet oak is also prominent in
several variants of the white oak (Q. alba)-black oak (Q.
velutina)-northern red oak (Q. rubra) SAF cover type [14].
At middle and lower elevations in the Appalachian Mountains, scarlet oak
is often a major component of pine (Pinus spp.) forests and pine heaths
[61]. Scarlet oak constitutes an important component of the subcanopy
and canopy layers of Table Mountain pine (Pinus pungens) forest [62].
The following published classifications list scarlet oak as a codominant
species:
Vegetation of the Great Smoky Mountains [61]
Old growth forests within the Piedmont of South Carolina [25]
MANAGEMENT CONSIDERATIONS
SPECIES: Quercus coccinea
WOOD PRODUCTS VALUE :
Although scarlet oak wood is of inferior grade, it is cut and utilized
with other red oaks as red oak lumber [20].
IMPORTANCE TO LIVESTOCK AND WILDLIFE :
Scarlet oak acorns are an important food source for numerous upland
wildlife species including squirrels, chipmunks, mice, wild turkeys,
white-tailed deer, blue jays, and woodpeckers [24]. White-tailed deer
occasionally browse young oak sprouts. The deer only take the top few
inches of the sprout unless it is extremely succulent or other food is
scarce [33].
Small mammals and birds use scarlet oak for nesting sites, both in the
canopy and in cavities [2,56].
PALATABILITY :
The sprouts of scarlet oak are more palatable to white-tailed deer than
the sprouts of bear oak [34].
NUTRITIONAL VALUE :
Scarlet oak acorns are on average 14.6 percent crude fat, 35.6 percent
total carbohydrates, 4.2 percent total protein, 0.18 percent calcium,
0.07 percent phosphorus, and 0.07 percent magnesium [4].
COVER VALUE :
NO-ENTRY
VALUE FOR REHABILITATION OF DISTURBED SITES :
NO-ENTRY
OTHER USES AND VALUES :
Scarlet oak is widely planted in the United States and Europe as a shade
tree and ornamental. It has brilliant red foliage in autumn [24].
OTHER MANAGEMENT CONSIDERATIONS :
Forest managers have noticed a decrease in upland oak frequency in newly
regenerated stands after clearcutting, especially on good sites. The
reason for the decrease is the inability of oak seedlings to compete
successfully with late successional, fast-growing species that have
invaded the oak forest understory in the absence of fire [51].
Oak seedlings that are occasionally top-killed will sprout from the
stump. These sprouts, known as advance regeneration, have
well-developed root systems. They grow faster than true seedlings and
are better able to compete successfully. To regenerate upland oaks
successfully, advance regeneration must be 4 to 5 feet (1.2-1.5 m) tall
before the overstory is removed. Regeneration of a mixed oak forest
after clearcutting can be successful only if there are adequate numbers
of older advance regeneration or saplings [51]. Sanders [50] recommends
there be at least 433 well-distributed oak sprouts and saplings per acre
(1,070/ha). Otherwise, a shelterwood silviculture system is needed to
give oak regeneration time and partial light to grow [24,51,59]. For
best results, the shelterwood cut should leave a 60 to 70 percent
stocking density. All nonoak stems in the understory larger than 4 to 6
feet (1.2-1.8 m) tall should be killed [51].
The season of clearcutting appears to have an effect on the regeneration
of upland oaks stands. On medium quality sites in south-central Ohio,
upland oaks (chestnut, scarlet, black and white) were more favored over
mixed hardwoods after summer clearcutting than after winter
clearcutting. The season of harvest (dormant season versus growing
season) did not affect regeneration on good sites [59].
Site quality affects the ability of upland oaks to regenerate. In the
above study in south-central Ohio, medium quality sites had higher
absolute and relative oak densities 18 to 20 years after clearcutting
than did good sites. The oaks showed good early establishment on both
medium and good sites but were later unable to compete with the faster
growing, mesic hardwoods on good sites [59].
Information on storage, seeding, and planting techniques for upland oaks
is detailed [49]. A method for increasing the growth rate of northern
red oak seedlings in nurseries has been developed and may be applicable
to scarlet oak [55].
Once scarlet oak stands are established, thinning increases the growth
rate of remaining trees. Thirty-two-year-old scarlet oaks showed 12
years of increased differential diameter growth beginning 6 to 7 years
after thinning. The reason for the delayed response is unknown [10].
Information on thinning, stocking, growth and yields of upland oaks is
detailed [18]. Thinning upland oak stands to retain the best acorn
producers for wildlife habitat enhancement did not improve acorn yields
enough to justify the efforts [13].
Scarlet oak is susceptible to a number of insects and diseases. Gypsy
moth (Lymantria dispar), an introduced species, defoliates scarlet oak,
and successive defoliations can kill a tree. Other insects that
defoliate scarlet oak include oak leaftier (Croesia semipurpurana), fall
cankerworm (Alsophila pometaria), forest tent caterpillar (Malacosoma
disstria), and walkingstick (Diapheromera femorata). Insects that bore
into the trunk include twolined chestnut borer (Agrilus bilineatus), red
oak borer (Enaphalodes rufulus), oak timberworm (Arrhenodes minutus),
Ambrosia beetles (Platypus spp. and Xyleborus spp.), and the larvae of
carpenterworms (Prionoxystus spp.). Egg-laying activity of the gouty
oak gall wasp (Callirhytis quercuepuntata) results in galls on smaller
twigs and limbs, and the oak-apple gall wasp (Amphibolips confluenta)
causes gall growth on leaves and petioles. Black carpenter ants
(Camponotus pennsylvanicus) sometimes nest in standing trees [24].
Scarlet oak is susceptible to oak wilt caused by the fungus Cerotocystis
fagacearum, and infected trees may die within 1 month. Cankers are
caused by Nectria spp. and Strummella coryneoidea. Fungi, such as
Stereum gausapatum, cause heart rot and enter the bole through branch
stubs and fire wounds. Scarlet oak does not self-prune well, and old
branch stubs facilitate fungi entry [24]. Pruning a scarlet oak stand
results in better timber quality [7]. Sprouts are susceptible to heart
rot; sprouts from large diameter stumps are more susceptible than those
from small diameter stumps [24].
Scarlet oaks that are stressed from drought, gypsy moth defoliation,
spring frost defoliation, old age, fire, poor site conditions, or other
factors often succumb to secondary agents such as twolined chestnut
borer and shoestring root rot (Armillaria mellea). This scenario, in
which a primary agent stresses the tree and a secondary agent kills it,
is known as "oak decline" and is responsible for considerable scarlet
oak mortality. For instance, from 1968 to 1972, 27 percent of scarlet
oak in the Newark Watershed in New Jersey died from gypsy moth
defoliation followed by twolined chestnut borer and shoestring root rot
attack [42]. Based on site factors, a general stand classification of
mortality risk from oak decline has been developed [53].
Herbicides have been used to control scarlet oak on sites where pine
regeneration is desired. In order to convert a North Carolina
Appalachian site to white pine (Pinus strobus), picloram was applied in
May as 10 percent acid equivalent pellets at the rate of 4.5 pounds acid
equivalent per acre (5.0 kg ae/ha). One year later, 19 percent of the
scarlet oaks showed complete crown kill or defoliation; 72 percent showed
leaf curling, crown biomass reduction, and/or chlorosis; and 9 percent
exhibited no effect from the herbicide treatment [44].
In Georgia, three herbicides were tested on scarlet oak. Each tree
received one incision for every 3 inches (7.6 cm) in d.b.h.; each
incision was injected with 0.06 ounces (2 ml) of herbicide. One year
after injection, scarlet oaks injected with Arsenal at two different
concentrations (1 and 2 pounds AC 252,925 per gallon [120 and 240 g/l])
had 100 percent top-kill and no sprouting. Garlon 3A (1.5 pounds
triclopyr per gallon [180 g/l]) resulted in 50 percent top-kill.
Scarlet oak injected with 3,6-dichloropicolinic acid at two
concentrations (1.5 and 3 pounds XRM-3972 per gallon [180 and 360 g/l])
resulted in 0 percent and 20 percent top-kill, respectively [41].
When managing forests for cavity-nesting species, scarlet oak should be
selected over hickories (Carya spp.) and white oak because of its high
number of cavities. In southeastern Missouri, 21.2 percent of scarlet
oaks had cavities. The average d.b.h. of cavity trees was 12.2 inches
(31.0 cm) with bigger trees having larger cavities [2]. In the Great
Smoky Mountains National Park, dead standing scarlet oak decays at a
rate of 5.7 percent per year [21].
BOTANICAL AND ECOLOGICAL CHARACTERISTICS
SPECIES: Quercus coccinea
GENERAL BOTANICAL CHARACTERISTICS :
Scarlet oak is a medium-sized, monoecious, native, deciduous tree with
an open, rounded crown [24,43]. At maturity, scarlet oak is usually 60
to 80 feet (18-24 m) tall and 24 to 36 inches (61-91 cm) in d.b.h., but
it can reach a maximum size of 100 feet (30 m) in height and 48 inches
(122 cm) in d.b.h. on good sites. Seedlings have a strong taproot and
relatively few lateral roots. Scarlet oak is one of the fastest growing
upland oak species [24] and is short-lived [38].
RAUNKIAER LIFE FORM :
Phanerophyte
REGENERATION PROCESSES :
Sexual: Seed production begins when the tree is about 20 years old,
with maximum production occurring after 50 years of age. Seed
production is irregular and unpredictable, but good crops generally
occur every 3 to 5 years [24]. Seeds are disseminated by animals and
gravity.
Germination is hypogeal. A light covering of forest litter is ideal for
germination [24]. In oak-pine forests in the New Jersey Pine Barrens,
scarlet oak seedlings occurred more frequently in areas with moss-lichen
cover and shallow litter layers than in areas with deep litter. The
mean litter depth of 1-year-old seedlings was 0.9 inch (2.2 cm) [7]. In
another study, the density of scarlet oak seedlings was negatively
correlated with deep litter coverage on some sites. However, this
negative correlation may be related to light availability because more
litter accumulated near shrubs [39].
A moderately open overstory canopy provides a favorable environment for
germination [24]. In the Pine Barrens of New Jersey, scarlet oak
seedlings occurred on sites with a higher percent (37.6) of full sun
than did chestnut oak or white oak. In addition, scarlet oak seedlings
occurred on sites with less competition, determined by distance to
closest neighbor in the ground layer. However, because acorns have
large energy reserves, 1-year-old seedlings may not be highly site
specific [7].
Vegetative: If top-killed, scarlet oak sprouts from dormant buds at or
above the root crown. Nearly 100 percent of stumps smaller than 4
inches (10 cm) in d.b.h. sprout, and about 18 percent of trees 24 inches
(61 cm) in d.b.h. sprout [24]. In a study on the Virginia Piedmont, the
season of harvest did not affect the the sprouting frequency of scarlet
oak stumps [27].
Scarlet oak stumps initially produce large numbers of sprouts [24], but
over time, sprout clumps tend towards the survival of one or two stems.
In one study, 5, 10, 15, 20, 25, and 35 years after cutting, the average
number of sprouts per stump was 9.0, 4.6, 2.7, 1.9, 1.5, and 1.3,
respectively [40]. Between the ages 4 and 8, stem-to-stem competition
within a scarlet oak clump is a more dominating interaction than
competition between clumps [8].
Scarlet oak sprouts grow faster in the first 5 years than the sprouts of
most associated oak species, but height growth falls off rapidly after
20 years [24]. The sprouts concentrate on stem growth during the first
and second growth flushes. A third flush does not show nearly as much
growth which may be because this species is adapted to xeric sites where
moisture stress limits growth later in the growing season [57].
Juveniles often die back and sprout numerous times, thus becoming
advance regeneration. Seedling sprouts grow faster than seedlings, with
the sprout growth rate dependent on the thickness of the stem [24].
SITE CHARACTERISTICS :
Scarlet oak, an upland xerophytic species, commonly occurs on ridges and
slopes in hilly to mountainous terrain. It occurs up to 5,000 feet
(1,520 m) in the southern Appalachian Mountains but is most common below
3,000 feet (910 m) [24]. In the Smoky Mountains, scarlet oak is most
frequent in middle and lower slope positions centered around 2,500 feet
(760 m) in elevation [61]. Scarlet oak will also grow in valley sites
on generally coarser soils than white oak [46].
Scarlet oak grows in a wide variety of soils, but especially in dry
sandy or gravelly soils [20,24]. It is most common on lower quality
sites [56]. In 51 upland hardwood stands on the Virginia Piedmont,
scarlet oak was significantly associated with low soil clay content
(p<0.05), low soil calcium (p<0.01), low soil magnesium (p<0.05), and
low pH (p<0.05) [15].
Common small tree and shrub associates of scarlet oak not mentioned in
Distribution and Occurrence include sassafras (Sassafras albidum),
flowering dogwood (Cornus florida), redbud (Cercis canadensis), sourwood
(Oxydendrum arboreum), sumacs (Rhus spp.), hawthorns (Crataegus spp.),
eastern hophornbeam (Ostrya virginiana), greenbriers (Smilax spp.),
blueberries (Vaccinium spp.), and huckleberries (Gaylussacia spp.).
Mountain-laurel (Kalmia latifolia) is an associate on very dry sites
[24,56].
SUCCESSIONAL STATUS :
Scarlet oak is intolerant of shade [24]. Seeds can germinate in the
shade, but seedlings do not exhibit long-term survival or growth under a
closed canopy [11]. Scarlet oak is usually found in dominant and
codominant positions, since suppressed individuals eventually die [24].
Scarlet oak tends to be better represented in forests with a history of
disturbance such as fire, logging, grazing, or disease [24,38]. In a
study of long-term forest composition in North Carolina, scarlet oak
regeneration was low for over 30 years, suggesting population
recruitment of this species is episodic and probably dependent on
disturbance. In the absence of disturbance, codominant scarlet oak
declines in importance in mixed oak stands [47].
Scarlet oak may be climax on dry sites with adequate light because of
its drought tolerance [24]. Little [32] suggests mixed oak forests of
black, white, chestnut, and scarlet oaks may represent a physiographic
climax association on upland sites in the New Jersey Pine Barrens.
SEASONAL DEVELOPMENT :
Scarlet oak flowers in April or May, depending on latitude, elevation,
and weather. Acorns mature in two growing seasons. They ripen and drop
in the fall and germinate in the spring [24].
FIRE ECOLOGY
SPECIES: Quercus coccinea
FIRE ECOLOGY OR ADAPTATIONS :
The fire resistance of scarlet oak is rated as low. It has thin bark,
and even low severity surface fires can result in severe basal damage
and high mortality. Top-killed scarlet oaks sprout vigorously from the
root crown after fire [24].
A thick litter covering is unfavorable for scarlet oak acorn germination
[24]. Fire removes litter and may facilitate scarlet oak regeneration
[39]. However, no study documenting increased scarlet oak regeneration
from seed after fire was found in the literature. The primary mode of
regeneration after fire appears to be sprouting.
The prevalent presettlement upland oak forests in the eastern and
central United States were associated with recurring fire. These
forests probably burned at an intermediate frequency (50 to 100 year
intervals), which promoted the dominance and stability of oak [1].
Oak-hickory forests are not usually conducive to high-severity fires,
but surface fires are easily ignited and spread rapidly under favorable
conditions [9]. In the absence of fire or other disturbance, the
short-lived scarlet oak is replaced by later successional species.
Periodic fire opens the canopy and provides an opportunity for upland
oaks to regenerate and maintain dominance [1]. Fifty-five years after a
late summer fire in south-central Connecticut, a burned area had higher
absolute and relative amounts of oak (scarlet, black, white, chestnut,
and northern red) than an adjacent unburned area [60].
In Missouri, nearly all trees were top-killed after a spring fire in a
23-year-old white oak-black oak-hickory stand in which 79 percent of the
stems larger than 0.6 inches (1.5 cm) in d.b.h. were oak and hickory.
Ten years later, 64 percent of the stems were oak and hickory,
indicating that a stand can replace itself. Scarlet oak increased from
a prefire density of 253 stems per acre (625 stems/ha) to a postfire
density of 329 stems per acre (813 stems/ha). Scarlet oak basal area
decreased from (2.7 m sq/ha) prefire to (0.6 m sq/ha) postfire.
However, scarlet oak maintained its position as the third most frequent
overstory species behind white oak and black oak. New stems were
primarily from basal sprouting [37].
Scarlet oak is restricted from the pine-scrub oak communities of the New
Jersey Pine Barrens because it does not produce viable seed at a young
enough age to become established in areas that burn every 8 to 12 years
[31,32]. In the Pine Barrens, scarlet oak is usually restricted to the
later successional forests along with white, black, and chestnut oaks
[19].
FIRE REGIMES :
Find fire regime information for the plant communities in which this
species may occur by entering the species name in the FEIS home page under
"Find Fire Regimes".
POSTFIRE REGENERATION STRATEGY :
Tree with adventitious-bud root crown/root sucker
Secondary colonizer - off-site seed
FIRE EFFECTS
SPECIES: Quercus coccinea
IMMEDIATE FIRE EFFECT ON PLANT :
Small scarlet oak are easily top-killed by low-severity surface fires,
and larger scarlet oak may suffer severe basal damage [24]. Scarlet oak
is less resistant to basal injury than black oak, white oak, or chestnut
oak. Scarlet oak is often severely wounded even when the area of
discolored bark is comparatively small. Fire wounds often extend far
beyond the region of bark discoloration [45].
Because bark thickens with age, the longer the fire interval, the
greater the chance a thin-barked species will develop bark thick enough
to protect it from a low-severity surface fire. Scarlet oak is reduced
in number by short-interval fires because of its thin bark and slow
growth [22].
Almost all scarlet oak in a 23-year-old mixed oak-hickory stand in
Missouri were top-killed by a spring fire [37].
Acorns cannot withstand the amount of heat usually generated in leaf
litter fires [28]. Acorns buried in the soil by animals may survive,
although documentation of this possibility with respect to scarlet oak
was not found in the literature.
DISCUSSION AND QUALIFICATION OF FIRE EFFECT :
NO-ENTRY
PLANT RESPONSE TO FIRE :
Scarlet oak individuals, including seedlings, sprout from the root crown
when top-killed. Individuals that survive fire often have basal fire
wounds.
Fire-wounded scarlet oaks sometime suffer a reduction in the rate of
diameter growth. A severe fire in the Bent Creek Experimental Forest in
North Carolina resulted in larger reductions in growth than a
moderate-severity fire. While unwounded scarlet oaks grew on average of
0.20 inch per year (0.50 cm/yr) in diameter, scarlet oak with 1 to 25
percent, 25 to 50 percent, and more than 50 percent of the base wounded
grew 0.17 inch per year (0.43 cm/yr), 0.14 inch per year (0.36 cm/yr),
and 0.09 inch per year (0.23 cm/y), respectively [23].
The mortality of oak trees from fire is often delayed. Six months after
two surface fires of different severity in southern New York, living
butt-scorched trees (larger than 1 inch [2.5 cm] in d.b.h.) were tagged
for future study. In the less severely burned area, 23 percent of the
tagged scarlet oak were dead 1.5 years after the fire. Smaller diameter
trees, especially those less than 5 inches (12.7 cm), had the highest
mortality. In the other area that burned more severely because of a
dense understory of mountain-laurel, 100 percent of the tagged scarlet
oaks were dead 1.5 years after the fire. The authors concluded that at
least one postfire growing season must elapse before fire damage to oaks
can be accurately determined [54].
The density of scarlet oak stems generally increases after fire because
of sprouting. Two growing seasons after two annual fires in an oak-pine
stand in the Cumberland Plateau in Kentucky, scarlet oak and black oak
stems increased from a prefire density of approximately 1,250 stems per
acre (3,090 stems/ha) to a postfire density of approximately 1,750 stems
per acre (4,320 stems/ha) [63].
If high fire frequency is continued, however, scarlet oak density will
eventually decrease as rootstocks weaken and die. After 27 years of
annual burning in a Tennessee upland oak forest, both overstory and
understory stem densities of scarlet oak were considerably reduced [11].
DISCUSSION AND QUALIFICATION OF PLANT RESPONSE :
The following Research Project Summaries
provide information on prescribed
fire use and postfire response of plant
community species, including scarlet
oak, that was not available when this
species review was originally
written:
FIRE MANAGEMENT CONSIDERATIONS :
Prescribed fire is often used to control hardwoods and promote
establishment of pine. In a study on the South Carolina Piedmont,
spring felling of leafed-out residual oaks (scarlet, chestnut, and
black) followed by summer burning produced greater reductions of
dominant sprout height and sprout clump crown diameters at the end of
the first postfire growing season than did winter felling followed by
summer broadcast burning. Spring felling was probably more effective
because carbohydrate root reserves are low after leaves emerge [17].
Because of the prolific sprouting of scarlet oak, prescribed burning
without additional use of herbicides or mechanical removal may not
adequately control this species. Prescribed burning is not recommended
for hardwood control, including scarlet oak, for shortleaf pine (Pinus
echinata) regeneration on the Cumberland Plateau in Kentucky [63].
Equations have been developed to estimate the fire-caused mortality of
scarlet oak. In order to predict mortality, a manager needs to know the
tree d.b.h., the height of bark blackening, the width of bark blackening
1 foot (0.3 m) above the ground, and the season of fire. The equation
should be applied to trees between 3 and 16 inches (7.6-40.6 cm) in
d.b.h. [35]. Equations have also been developed to predict lumber value
losses due to fire wounding of scarlet oak [36]. An equation has been
developed to predict the size of a fire wound on a scarlet oak from the
area of the exterior discolored bark and the diameter of the damaged
tree [45].
Scarlet oak is a preferred species for shelterbelts around farms and
houses to protect them from fire in open country. The row of trees
reduces wind velocity, filters out air borne debris, slows ground fires,
and readily regenerates after fire [52].
REFERENCES
SPECIES: Quercus coccinea
REFERENCES :
1. Abrams, Marc D. 1992. Fire and the development of oak forests.
BioScience. 42(5): 346-353. [19215]
2. Allen, Arthur W.; Corn, Janelle G. 1990. Relationships between live tree
diameter and cavity abundance in a Missouri oak-hickory forest. Northern
Journal of Applied Forestry. 7: 179-183. [13504]
3. Bonner, F. T.; Vozzo, J. A. 1987. Seed biology and technology of
Quercus. Gen. Tech. Rep. SO-66. New Orleans, LA: U.S. Department of
Agriculture, Forest Service, Southern Forest Experiment Station. 21 p.
[3248]
4. Braun, E. Lucy. 1961. The woody plants of Ohio. Columbus, OH: Ohio State
University Press. 362 p. [12914]
5. Brown, Arthur A.; Davis, Kenneth P. 1973. Forest fire control and use.
2nd ed. New York: McGraw-Hill. 686 p. [15993]
6. Burns, Paul Y.; Nichols, J. Milford. 1952. Oak pruning in the Missouri
Ozarks. University of Missouri Agricultural Experiment Station Bulletin.
581(Apr): 1-8. [10156]
7. Collins, Scott L.; Good, Ralph E. 1987. The seedling regeneration niche:
habitat structure of tree seedlings in an oak-pine forest. Oikos. 48:
89-98. [8637]
8. Cook, James E. 1990. Degree of competition and integration in one- to
eight-year-old scarlet and chestnut oak sprout clumps. In: Van Sambeek,
J. W.; Larson, M. M., eds. Proceedings, 4th workshop on seedling
physiology and growth problems in oak plantings; 1989 March 1-2;
Columbus, OH. (Abstracts). Gen. Tech. Rep. NC-139. St. Paul, MN: U.S.
Department of Agriculture, Forest Service, North Central Forest
Experiment Station: 29. Abstract. [13156]
9. Crosby, John S.; Loomis, Robert M. 1974. Some forest floor fuelbed
characteristics of black oak stands in southeast Missouri. NC-162. St.
Paul, MN: U.S. Department of Agriculture, Forest Service, North Central
Forest Experiment Station. 4 p. [8153]
10. Cutter, Bruce E.; Lowell, Kim E.; Dwyer, John P. 1991. Thinning effects
on diameter growth in black and scarlet oak as shown by tree ring
analyses. Forest Ecology and Management. 43: 1-13. [16684]
11. DeSelm, Hal R.; Clebsch, Edward E. C.; Rennie, John C. 1991. Effects of
27 years of prescribed fire on an oak forest and its soils in middle
Tennessee. In: Coleman, Sandra S.; Neary, Daniel G., compiler.
Proceedings, 6th biennial southern silvicultural research conference:
Vol. 1; 1990 October 30 - November 1; Memphis, TN. Gen. Tech. Rep.
SE-70. Asheville, NC: U.S. Department of Agriculture, Forest Service,
Southeastern Forest Experiment Station: 409-417. [17488]
12. Dibble, Alison C.; Campbell, Christopher S.; Tyler, Harry R., Jr.;
Vickery, Barbara St. J. 1989. Maine's official list of endangered and
threatened plants. Rhodora. 91(867): 244-269. [4258]
13. Drake, William E. 1991. Evaluation of an approach to improve acorn
production during thinning. In: McCormick, Larry H.; Gottschalk, Kurt
W., eds. Proceedings, 8th central hardwood forest conference; 1991 March
4-6; University Park, PA. Gen. Tech. Rep. NE-148. Radnor, PA: U.S.
Department of Agriculture, Forest Service, Northeastern Forest
Experiment Station: 429-441. [15328]
14. Eyre, F. H., ed. 1980. Forest cover types of the United States and
Canada. Washington, DC: Society of American Foresters. 148 p. [905]
15. Farrell, John D.; Ware, Stewart. 1991. Edaphic factors and forest
vegetation in the piedmont of Virgina. Bulletin of the Torrey Botanical
Club. 118(2): 161-169. [15694]
16. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others].
1977. Vegetation and environmental features of forest and range
ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of
Agriculture, Forest Service. 68 p. [998]
17. Geisinger, Donn R.; Waldrop, Thomas A.; Haymond, Jacqueline L.; Van
Lear, David H. 1989. Sprout growth following winter and spring felling
with and without summer broadcast burning. In: Waldrop, Thomas A., ed.
Proceedings of pine-hardwood mixtures: a symposium on management and
ecology of the type; 1989 April 18-19; Atlanta, GA. Gen. Tech. Rep.
SE-58. Asheville, SC: U.S. Department of Agriculture, Forest Service,
Southeastern Forest Experiment Station: 91-95. [10262]
18. Gingrich, Samuel F. 1971. Stocking, growth, and yield of oak stands. In:
Oak symposium: Proceedings; 1971 August 16-20; Morgantown, WV. Upper
Darby, PA: U.S. Department of Agriculture, Forest Service, Northeastern
Forest Experiment Station: 65-73. [9085]
19. Givnish, Thomas J. 1981. Serotiny, geography, and fire in the pine
barrens of New Jersey. Evolution. 35(1): 101-123. [8634]
20. Harlow, William M.; Harrar, Ellwood S., White, F. M. 1979. Textbook of
dendrology. 6th ed. New York: McGraw-Hill, Inc. 510 p. [18070]
21. Harmon, Mark E. 1982. Decomposition of standing dead trees in the
southern Appalachian Mountains. Oecologia. 52: 214-215. [13735]
22. Harmon, Mark E. 1984. Survival of trees after low-intensity surface
fires in Great Smoky Mountains National Park. Ecology. 65(3): 796-802.
[10997]
23. Jemison, George M. 1944. The effect of basal wounding by forest fires on
the diameter growth of some southern appalachian hardwoods. Bulletin 9.
Durham, NC: Duke University, School of Forestry. 63 p. [8716]
24. Johnson, Paul S. 1990. Quercus coccinea Muenchh. scarlet oak. In:
Burns, Russell M.; Honkala, Barbara H., tech. coords. Silvics of North
America. Vol. 2, Hardwoods. Agric. Handb. 654. Washington, DC: U.S.
Department of Agriculture, Forest Service: 625-630. [19373]
25. Jones, Steven M. 1988. Old-growth forests within the Piedmont of South
Carolina. Natural Areas Journal. 8(1): 31-37. [11008]
26. Kartesz, John T.; Kartesz, Rosemarie. 1980. A synonymized checklist of
the vascular flora of the United States, Canada, and Greenland. Volume
II: The biota of North America. Chapel Hill, NC: The University of North
Carolina Press; in confederation with Anne H. Lindsey and C. Richie
Bell, North Carolina Botanical Garden. 500 p. [6954]
27. Dieterich, John H. 1983. Fire history of southwestern mixed conifer: a
case study. Forest Ecology. 6: 13-31. [5242]
28. Korstian, C. F. 1927. Factors controlling germination and early survival
of oaks. Bull. No. 19. New Haven, CT: Yale University, School of
Forestry. 115 p. [19369]
29. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation
of the conterminous United States. Special Publication No. 36. New York:
American Geographical Society. 77 p. [1384]
30. Little, Elbert L., Jr. 1979. Checklist of United States trees (native
and naturalized). Agric. Handb. 541. Washington, DC: U.S. Department of
Agriculture, Forest Service. 375 p. [2952]
31. Little, S. 1964. Fire ecology and forest management in the New Jersey
pine region. In: Proceedings, 3rd annual Tall Timbers fire ecology
conference; 1964 April 9-10; Tallahassee, FL. No. 3. Tallahassee, FL:
Tall Timbers Research Station: 35-59. [5893]
32. Martin, S. Clark. 1980. Mesquite. In: Eyre, F. H., ed. Forest cover
types of the United States and Canada. Washington, DC: Society of
American Foresters: 118. [9858]
33. Little, Silas; Moorhead, George R.; Somes, Horace A. 1958. Forestry and
deer in the Pine Region of New Jersey. Station Pap. No. 109. Upper
Darby, PA: U.S. Department of Agriculture, Forest Service, Northeastern
Forest Experiment Station. 33 p. [11681]
34. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession
following large northern Rocky Mountain wildfires. In: Proceedings, Tall
Timbers fire ecology conference and Intermountain Fire Research Council
fire and land management symposium; 1974 October 8-10; Missoula, MT. No.
14. Tallahassee, FL: Tall Timbers Research Station: 355-373. [1496]
35. Loomis, Robert M. 1973. Estimating fire-caused mortality and injury in
oak-hickory forests. Res. Pap. NC-94. St. Paul, MN: U.S. Department of
Agriculture, Forest Service, North Central Forest Experiment Station. 6
p. [8740]
36. Loomis, Robert M. 1974. Predicting the losses in sawtimber volume and
quality from fires in oak-hickory forests. NC-104. St. Paul, MN: U.S.
Department of Agriculture, Forest Service, North Central Forest
Experiment Station. 6 p. [8712]
37. Loomis, Robert M. 1977. Wildfire effects on an oak-hickory forest in
southeast Missouri. Res. Note NC-219. St. Paul, MN: U.S. Department of
Agriculture, Forest Service, North Central Forest Experiment Station. 4
p. [8738]
38. Martin, William H. 1992. Characteristics of old-growth mesophytic
forests. Natural Areas Journal. 12(3): 127-135. [19371]
39. Matlack, G. R.; Good, R. E. 1989. Plant-scale pattern among herbs and
shrubs of a fire-dominated coastal plain forest. Vegetatio. 82: 95-103.
[9829]
40. McIntyre, A. C. 1936. Sprout groups and their relation to the oak
forests of Pennsylvania. Journal of Forestry. 34: 1054-1058. [10086]
41. Michael, J. L. 1985. Hardwood control by injection with two new
chemicals. Proceedings of the Southern Weed Science Society. 38:
164-167. [12687]
42. Millers, Imants; Shriner, David S.; Rizzo, David. 1989. History of
hardwood decline in the eastern United States. Gen. Tech. Rep. NE-126.
Bromall, PA: U.S. Department of Agriculture, Forest Service,
Northeastern Forest Experiment Station. 75 p. [10925]
43. National Arbor Day Foundation. 1992. Scarlet oak. Arbor Day. Sept/Oct:
8. [19370]
44. Neary, D. G.; Douglass, J. E.; Ruehle, J. L.; Fox, W. 1984. Converting
rhododendron-laurel thickets to white pine with picloram and
mycorrhizae-inoculated seedlings. Southern Journal of Applied Forestry.
8(3): 163-168. [10697]
45. Nelson, Ralph M.; Sims, Ivan H.; Abell, Margaret S. 1933. Basal fire
wounds on some southern Appalachian hardwoods. Journal of Forestry. 31:
829-837. [160]
46. Nowacki, Gregory J.; Abrams, Marc D. 1991. Community and edaphic
analysis of mixed oak forests in the Ridge and Valley Province of
central Pennsylvania. In: McCormick, Larry H.; Gottschalk, Kurt W., eds.
Proceedings, 8th central hardwood forest conference; 1991 March 4-6;
University Park, PA. Gen. Tech. Rep. NE-148. Radnor, PA: U.S. Department
of Agriculture, Forest Service, Northeastern Forest Experiment Station:
247-260. [15315]
47. Phillips, Donald L.; Murdy, William H. 1985. Effects of Rhododendron
(Rhododendron maximum L.) on regeneration of southern Appalachian
hardwoods. Forest Science. 31(1): 226-233. [19372]
48. Raunkiaer, C. 1934. The life forms of plants and statistical plant
geography. Oxford: Clarendon Press. 632 p. [2843]
49. Russell, T. E. 1971. Seeding and planting upland oaks. In: Oak
symposium: Proceedings; 1971 August 16-20; Morgantown, WV. Upper Darby,
PA: U.S. Department of Agriculture, Forest Service, Northeastern Forest
Experiment Station: 49-54. [9082]
50. Sander, Ivan L. 1977. Manager's handbook for oaks in the North Central
States. Gen. Tech. Rep NC-37. St. Paul, MN: U.S. Department of
Agriculture, Forest Service, North Central Forest Experiment Station. 35
p. [11002]
51. Sander, Ivan L. 1988. Guidelines for regenerating Appalachian oak
stands. In: Smith, H. Clay; Perkey, Arlyn W.; Kidd, William E., Jr.,
eds. Guidelines for regenerating Appalachian hardwood stands: Workshop
proceedings; 1988 May 24-26; Morgantown, WV. SAF Publ. 88-03.
Morgantown, WV: West Virginia University Books: 189-198. [13945]
52. Simpfendorfer, K. J. 1989. Trees, farms and fires. Land and Forests
Bulletin No. 30. Victoria, Australia: Department of Conservation,
Forests and Lands, Lands and Forests Division. 55 p. [10649]
53. Starkey, Dale A.; Oak, Steven W. 1989. Site factors and stand conditions
associated with oak decline in Southern upland hardwood forests. In:
Rink, George; Budelsky, Carl A., eds. Proceedings, 7th central hardwood
conference; 1989 March 5-8; Carbondale, IL. Gen. Tech. Rep. NC-132. St.
Paul, MN: U.S. Department of Agriculture, Forest Service, North Central
Forest Experiment Station: 95-102. [9372]
54. Stickel, Paul W. 1935. Forest fire damage studies in the Northeast. II.
First-year mortality in burned-over oak stands. Journal of Forestry. 33:
595-598. [18764]
55. Struve, Daniel K. 1990. Red oak whip production in containers. In: Van
Sambeek, J. W.; Larson, M. M., eds. Proceedings, 4th workshop on
seedling physiology and growth problems in oak plantings; 1989 March
1-2; Columbus, OH. (Abstracts). Gen. Tech. Rep. NC-139. St. Paul, MN:
U.S. Department of Agriculture, Forest Service, North Central Forest
Experiment Station: 17. Abstract. [13144]
56. Trimble, George R., Jr.; Patric, James H.; Gill, John D.; [and others].
1974. Some options for managing forest land in the central Appalachians.
Gen. Tech. Rep. NE-12. Upper Darby, PA: U.S. Department of Agriculture,
Forest Service, Northeastern Forest Experiment Station. 42 p. [13545]
57. Tworkoski, T. J.; Ross, M. S.; Hopper, G. M. 1990. Analysis of chestnut
and scarlet oak stump growth. Canadian Journal of Forestry Research. 20:
112-116. [11125]
58. U.S. Department of Agriculture, Soil Conservation Service. 1982.
National list of scientific plant names. Vol. 1. List of plant names.
SCS-TP-159. Washington, DC. 416 p. [11573]
59. Ward, Jeffery S.; Heiligmann, Randall B. 1990. Effects of site quality
and season of clearcutting on hardwood regeneration in Ohio. Northern
Journal of Applied Forestry. 7: 69-72. [11879]
60. Ward, Jeffrey S.; Stephens, George R. 1989. Long-term effects of a 1932
surface fire on stand structure in a Connecticut mixed hardwood forest.
In: Rink, George; Budelsky, Carl A., eds. Proceedings, 7th central
hardwood conference; 1989 March 5-8; Carbondale, IL. Gen. Tech. Rep.
NC-132. St. Paul, MN: U.S. Department of Agriculture, Forest Service,
North Central Forest Experiment Station: 267-273. [9389]
61. Whittaker, R. H. 1956. Vegetation of the Great Smoky Mountains.
Ecological Monographs. 26(1): 1-79. [11108]
62. Williams, Charles E.; Johnson, W. Carter. 1990. Age structure and the
maintenance of Pinus pungens in pine-oak forests of southwestern
Virginia. American Midland Naturalist. 124(1): 130-141. [12747]
63. Williamson, Malcolm J. 1964. Burning does not control young hardwoods on
shortleaf pine sites in the Cumberland Plateau. Res. Note CS-19.
Columbus, OH: U.S. Department of Agriculture, Forest Service, Central
States Forest Experiment Station. 4 p. [10999]
FEIS Home Page
https://www.fs.usda.gov/database/feis/plants/tree/quecoc/all.html