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SPECIES:  Pinus contorta var. contorta

Introductory

SPECIES: Pinus contorta var. contorta
AUTHORSHIP AND CITATION : Cope, Amy B. 1993. Pinus contorta var. contorta. In: Fire Effects Information System, [Online]. U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: https://www.fs.usda.gov/database/feis/plants/tree/pinconc/all.html [].
ABBREVIATION : PINCONC PINCON SYNONYMS : Pinus boursieri Carr. Pinus tenuis Lemmon SCS PLANT CODE : PICO COMMON NAMES : shore pine beach pine coast pine lodgepole pine TAXONOMY : The currently accepted scientific name of shore pine is Pinus contorta Dougl. ex Loud var. contorta [24,33]. Shore pine is one of four varieties of lodgepole pine. The other three varieties are [9,24]: Rocky Mountain lodgepole pine (Pinus contorta var. latifolia), Sierra lodgepole pine (Pinus contorta var. murrayana), Mendocino White Plains lodgepole pine (Pinus contorta var. bolanderi). This write-up will focus on shore pine. LIFE FORM : Tree FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY

DISTRIBUTION AND OCCURRENCE

SPECIES: Pinus contorta var. contorta
GENERAL DISTRIBUTION : Shore pine occurs along the Pacific Coast from Yakutat Bay, Alaska, south through the Coast Ranges to Mendocino County, California [9,10,24,31,33]. In the eastern part of its range, shore pine occurs intermittently with Rocky Mountain lodgepole pine in the Cascade Range of northwestern Washington, British Columbia, and Alaska [38]. Shore pine is also found in the Klamath Mountains of Oregon and California [49]. ECOSYSTEMS : FRES23 Fir - spruce FRES24 Hemlock - Sitka spruce FRES26 Lodgepole pine FRES27 Redwood FRES28 Western hardwoods STATES : AK CA HI OR WA BC BLM PHYSIOGRAPHIC REGIONS : 1 Northern Pacific Border 3 Southern Pacific Border KUCHLER PLANT ASSOCIATIONS : K001 Spruce - cedar - hemlock forest K003 Silver fir - Douglas-fir forest K004 Fir - hemlock forest K006 Redwood forest K008 Lodgepole pine - subalpine forest K009 Pine - cypress forest K013 Cedar - hemlock - pine forest SAF COVER TYPES : 205 Mountain hemlock 206 Engelmann spruce - subalpine fir 215 Western white pine 218 Lodgepole pine 223 Sitka spruce 224 Western hemlock 225 Western hemlock - Sitka spruce 226 Coastal true fir - hemlock 227 Western redcedar - western hemlock 228 Western redcedar 232 Redwood 255 California coast live oak SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Shore pine is a dominant species in the northern part of its range. Farther south, shore pine is a codominant or subdominant species. In mixed stands, shore pine may form scrubby thickets or sparse to dense groves. Other vegetation is usually sparse [1,48]. Shore pine is listed as an indicator in the following published classifications: Provisional plant community types of southeastern Alaska [1] A classification system for California's hardwood rangelands [2] Preliminary forest plant association management guide [11] Preliminary forest plant associations of the Stikine Area, Tongass National Forest [42] The Alaska vegetation classification system [46] The closed-cone pine and cypresses [48].

MANAGEMENT CONSIDERATIONS

SPECIES: Pinus contorta var. contorta
WOOD PRODUCTS VALUE : The wood of shore pine is light, brittle, coarse grained, and has a high specific gravity [18,33]. Shore pine is occasionally used as fuel and produces 8,730 British thermal units per pound [18]. IMPORTANCE TO LIVESTOCK AND WILDLIFE : Rodents eat the seeds of shore pine [27], and porcupines consume the cambium [3]. Shore pine is of slight importance to big game but provides important edge habitat for other animals [6,42,44]. Alaskan brown bears travel through corridors of shore pine while traveling to feeding areas. Shore pine provides nesting habitat for yellowlegs in Alaska [11]. PALATABILITY : NO-ENTRY NUTRITIONAL VALUE : NO-ENTRY COVER VALUE : NO-ENTRY VALUE FOR REHABILITATION OF DISTURBED SITES : Shore pine survives strong, salty winds on dry crests and wet depressions. Shore pine has helped stabilize recent sand dune expansion in California [20]. OTHER USES AND VALUES : Native Americans boiled the inner bark of shore pine for food [16]. Coastal Native Americans used the pitch of shore pine to treat open sores and chewed the buds to relieve sore throats [3]. OTHER MANAGEMENT CONSIDERATIONS : Shore pine is a useful species for watershed stabilization [22]. It has shown potential in shelterbelt plantings because of its branching habit and winter hardiness [22,28]. Road construction should be avoided on shore pine sites where possible because of the exceptionally deep, wet soils [11]. Shore pine sites are important because they absorb excessive rainfall and regulate waterflows [11]. Shore pine is valued for the rapid early growth of seedlings [22]. Shore pine is a primary host to lodgepole pine dwarf mistletoe (Arceuthobium americanum); however, infection by this parasite has been observed only in a few coastal areas of British Columbia [14,21]. Shore pine is a host to hemlock dwarf mistletoe (Arceuthobium tsugense), which can cause localized infections [5,30].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Pinus contorta var. contorta
GENERAL BOTANICAL CHARACTERISTICS : Shore pine is a short-lived, native conifer [9,22,25,43]. At maturity, depending on the site, shore pine reaches heights of 20 to 50 feet (6-15 m) and d.b.h.'s of 6 to 20 inches (15-50 cm) [16,26,40,46]. The trunk is often twisted, and crown shape varies from dense and round to irregular [17,19,27]. The bark of shore pine is furrowed and up to 1 inch (2.54 cm) thick [3,9,26]. Shore pine has many branches [4,9,26]. The short, narrow leaves occur in fascicles of two [3,17,19,26,44]. The cones are persistent [19,22,33,44,49]. They are about 1.5 to 2 inches (3.8-5.1 cm) long [19,49]. Cones of shore pine are typically nonserotinous [9,10,26]. Serotiny tends to increase farther inland but is erratic and unpredictable [26]. RAUNKIAER LIFE FORM : Phanerophyte REGENERATION PROCESSES : Shore pine does not reproduce by sprouting [40]. Shore pine begins producing cones between 5 and 10 years of age [10,22]. Good seed crops usually occur every other year [10]. Owen and Molden [50] discuss development of lateral shoot terminal buds. Shore pine produces large amounts of flowers and pollen [3,9,49]. Percentage of sound seed ranges from 75 to 79 percent [9]. Fresh seed requires no stratification, but stored seed requires 20 to 30 days stratification. Seeds are viable for up to 17 years in cold storage [22]. Seed falls approximately 200 feet (60 m) from the source under normal conditions [9]. Shore pine requires absorbent soils, light, and warm temperatures for germination [8,22]. SITE CHARACTERISTICS : Shore pine grows in a maritime climate throughout most of its range. Annual precipitation, which falls year-round and mostly in the form of rain, is 60 to 200 inches (1,500-5,000 mm) [15,26]. Shore pine occurs from sea level to the subalpine zone (5,030 feet [1,525 m]) in Alaska [3,45]. Shore pine occurs in peat bogs and muskegs on gentle slopes and lowlands [11,19,40,42,49]. It is most common on poorly drained, deep Histosols [11,42,45]. Conditions are xeric in the southern-most part of shore pine's distribution, where it occurs in closed-cone pine and cypress communities of California [48]. Shore pine habitat here includes coastal dunes, seaside bluffs, and exposed rocky headlands; winds may be strong and salty [48]. Soils are Inceptisols, Alfisols, and Ultisols; best growth is on well-drained loams with a pH of 5 [16,26]. Here shore pine occurs at elevations between sea level and 1,690 feet (0-507 m) [33]. Overstory and understory tree species not mentioned in Distribution and Occurrence include yellow-cedar (Chamaecyparis nootkatensis), Mendocino White Plains lodgepole pine, bishop pine (Pinus muricata), and common juniper (Juniperus communis) [20,34,47]. Associated shrubs are huckleberries (Vaccinium spp.), bog Labrador-tea (Ledum groenlandicum), bog kalmia (Kalmia polifolia), northern twinflower (Linnaea borealis), bunchberry dogwood (Cornus canadensis), and crowberry (Empetrum nigrum) [1,8,41,46,47]. Commonly associated herbs are sedges (Carex spp.), naked sedge (Calamagrostis nutkatensis), rusty menziesia (Menziesia ferruginea), and narrowleaf cottonsedge (Eriophorum angustifolium) [8,46,47]. Sphagnum moss (Sphagnum spp.) is a common associate in Alaska [46,47]. SUCCESSIONAL STATUS : Facultative Seral Species Shore pine occurs in extreme habitats that are unfavorable to other, potentially competitive species [9,49]. Windthrow and landslides are common [15]. Shore pine is a climatic climax in bog woodlands, reproducing under its own canopy [36]. Shore pine is considered a climax species in many areas [6,41,46]. Shore pine is shade intolerant [25,26,40]. SEASONAL DEVELOPMENT : Male and female strobili are initiated late in the growing season. Development continues in the spring [26]. Pollen shedding begins in mid- to late May [9,10,26]. Cones mature between September and October, with seed dispersal following shortly afterward [22,26,40].

FIRE ECOLOGY

SPECIES: Pinus contorta var. contorta
FIRE ECOLOGY OR ADAPTATIONS : Fire is not an important factor in forest succession where shore pine occurs in southeastern Alaska [15]. Fire is infrequent in maritime forest types and usually is of little ecological significance [25]. The fire interval is 150 to 350 years but may not be cyclic [35]. The coastal cedar-pine-hemlock biogeoclimatic zone of British Columbia has little or no fire history. The presence of shade-tolerant firs and hemlocks and fire0sensitive species, such as shore pine, indicate that fire is rare in this area [34]. The coastal dunes where shore pine occurs in California are considered fire-free [48]. The foliage of shore pine is moderately flammable. Shore pine has a moderate to low degree of fire resistance [25]. FIRE REGIMES : Find fire regime information for the plant communities in which this species may occur by entering the species name in the FEIS home page under "Find Fire Regimes". POSTFIRE REGENERATION STRATEGY : Tree without adventitious-bud root crown Secondary colonizer - off-site seed

FIRE EFFECTS

SPECIES: Pinus contorta var. contorta
IMMEDIATE FIRE EFFECT ON PLANT : Shore pine is sensitive to fire and probably killed by most fires. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : NO-ENTRY DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : NO-ENTRY

REFERENCES

SPECIES: Pinus contorta var. contorta
REFERENCES : 1. Alaback, Paul B. 1980. Provisional plant community types of southeastern Alaska. Unpublished paper on file at: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Intermountain Fire Sciences Laboratory, Missoula, MT. 15 p. [18773] 2. Allen, Barbara H.; Holzman, Barbara A.; Evett, Rand R. 1991. A classification system for California's hardwood rangelands. Hilgardia. 59(2): 1-45. [17371] 3. Arno, Stephen F.; Hammerly, Ramona P. 1977. Northwest trees. Seattle, WA: The Mountaineers. 222 p. [4208] 4. Arno, Stephen F.; Hammerly, Ramona P. 1984. Timberline: Mountain and arctic forest frontiers. Seattle, WA: The Mountaineers. 304 p. [339] 5. Baranyay, J. A.; Smith, R. B. 1972. Dwarf mistletoes in British Columbia and recommendations for their control. BC-X-72. Victoria, BC: Canadian Forestry Service, Pacific Forest Research Centre. 18 p. [16391] 6. Bartolome, James W. 1983. Overstory-understory relationships: lodgepole pine forest. In: Bartlett, E. T.; Betters, David R., eds. Overstory-understory relationships in western forests. Western Regional Research Publication No. 1. Fort Collins, CO: Colorado State University Experiment Station: 1-4. [3308] 7. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 8. Coates, K. David. 1987. Effects of shrubs and herbs on conifer regeneration and microclimate in the Rhododendron-Vaccinium-Menziesia community of south-central BC. Vancouver, BC: University of British Columbia. Thesis. Abstract. [17445] 9. Critchfield, W. B. 1978. The distribution, genetics, and silvics of lodgepole pine. In: Proceedings of the IUFRO joint meeting of working parties, Volume one: background papers and Douglas fir provenances; [Date of conference unknown]; Vancouver, B.C., Canada. Volume one. Victoria, B.C., Canada: British Columbia Ministry of Forests: 65-94. [8317] 10. Critchfield, William B. 1980. Genetics of lodgepole pine. Res. Pap. WO-37. Washington, DC: U.S. Department of Agriculture, Forest Service. 57 p. [8283] 11. DeMeo, Thomas. 1989. Preliminary forest plant association management guide: Ketchikan Area, Tongass National Forest. [Portland, OR]: [U.S. Department of Agriculture, Forest Service]. 164 p. [19017] 12. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 13. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 14. Hawksworth, Frank G.; Johnson, David W. 1989. Biology and management of dwarf mistletoe in lodgepole pine in the Rocky Mountains. Gen. Tech. Rep. RM-169. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 38 p. [8651] 15. Arno, Stephen F.; Hoff, Raymond J. 1989. Silvics of whitebark pine (Pinus albicaulis). Gen. Tech. Rep. INT-253. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 11 p. [1304] 16. Hosie, R. C. 1969. Native trees of Canada. 7th ed. Ottawa, ON: Canadian Forestry Service, Department of Fisheries and Forestry. 380 p. [3375] 17. Hitchcock, C. Leo; Cronquist, Arthur. 1973. Flora of the Pacific Northwest. Seattle, WA: University of Washington Press. 730 p. [1168] 18. Howard, James O.; Setzer, Theodore S. 1989. Logging residue in southeast Alaska. Res. Pap. PNW-RP-405. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 36 p. [13189] 19. Hulten, Eric. 1968. Flora of Alaska and neighboring territories. Stanford, CA: Stanford University Press. 1008 p. [13403] 20. Jenny, H.; Arkley, R. J.; Schultz, A. M. 1969. The pygmy forest-podsol ecosystem and its dune associates of the Mendocino Coast. Madrono. 20: 60-74. [10726] 21. Kimmey, J. W. 1957. Dwarfmistletoes of California and their control. Tech. Pap. No. 19. Berkeley, CA: U.S. Department of Agriculture, Forest Service, California Forest and Range Experiment Station. 12 p. [16464] 22. Krugman, Stanley L.; Jenkinson, James L. 1974. Pinaceae--pine family. In: Schopmeyer, C. S., technical coordinator. Seeds of woody plants in the United States. Agric. Handb. 450. Washington, DC: U.S. Department of Agriculture, Forest Service: 598-637. [1380] 23. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 24. Little, Elbert L., Jr. 1979. Checklist of United States trees (native and naturalized). Agric. Handb. 541. Washington, DC: U.S. Department of Agriculture, Forest Service. 375 p. [2952] 25. Lotan, James E.; Alexander, Martin E.; Arno, Stephen F.; [and others]. 1981. Effects of fire on flora: A state-of-knowledge review. National fire effects workshop; 1978 April 10-14; Denver, CO. Gen. Tech. Rep. WO-16. Washington, DC: U.S. Department of Agriculture, Forest Service. 71 p. [1475] 26. Lotan, James E.; Critchfield, William B. 1990. Pinus contorta Dougl. ex. Loud. lodgepole pine. In: Burns, Russell M.; Honkala, Barbara H., technical coordinators. Silvics of North America. Volume 1. Conifers. Agric. Handb. 654. Washington, DC: U.S. Department of Agriculture, Forest Service: 302-315. [13393] 27. Lotan, James E.; Perry, David A. 1983. Ecology and regeneration of lodgepole pine. Agric. Handb. 606. Washington, DC: U.S. Department of Agriculture, Forest Service. 51 p. [8288] 28. Low, Alan J. 1986. Tree planting in the Falkland Islands. Forestry. 59(1): 59-84. [9755] 30. Mathiasen, Robert L.; Hawksworth, Frank G. 1988. Dwarf mistletoes on western white pine and whitebark pine in northern California and southern California. Forest Science. 34(2): 429-440. [5034] 31. McMillan, Calvin. 1956. The edaphic restriction of Cupressus and Pinus in the Coast Ranges of central California. Ecological Monographs. 26: 177-212. [11884] 32. Millar, Constance I.; Libby, William J. 1989. Disneyland or native ecosystem: genetics and the restorationist. Restoration and Management Notes. 7(1): 18-24. [8071] 33. Munz, Philip A. 1973. A California flora and supplement. Berkeley, CA: University of California Press. 1905 p. [6155] 34. Parminter, John. 1983. Fire history and fire ecology in the Prince Rupert Forest region. In: Trowbridge, R. L.; Macadam, A., eds. Prescribed fire--forest soils: Symposium proceedings; 1982 March 2-3; Smithers, BC. Land Management Report Number 16. Victoria, BC: Province of British Columbia, Ministry of Forests: 1-35. [8849] 35. Parminter, John. 1991. Fire history and effects on vegetation in three biogeoclimatic zones of British Columbia. In: Nodvin, Stephen C.; Waldrop, Thomas A., eds. Fire and the environment: ecological and cultural perspectives: Proceedings of an international symposium; 1990 March 20-24; Knoxville, TN. Gen. Tech. Rep. SE-69. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southeastern Forest Experiment Station: 263-272. [16651] 36. Pojar, Jim. 1985. Ecological classification of lodgepole pine in Canada. In: Baumgartner, David M.; Krebill, Richard G.; Arnott, James T.; Weetman, Gordon F., compilers and editors. Lodgepole pine: The species and its management: Symposium proceedings; 1984 May 8-10; Spokane, WA; 1984 May 14-16; Vancouver, BC. Pullman, WA: Washington State University, Cooperative Extension: 77-88. [9442] 37. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 38. Shrimpton, D. M. 1972. Variation in the extractives from lodgepole pine sapwood and heartwood. Information Report NOR-X-18. Edmonton, Alberta: Environment Canada, Forestry Service, Northern Forest Research Centre. 22 p. [8323] 39. Stickney, Peter F. 1989. Seral origin of species originating in northern Rocky Mountain forests. Unpublished draft on file at: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory, Missoula, MT; RWU 4403 files. 7 p. [20090] 40. Tackle, David. 1961. Silvics of lodgepole pine. Misc. Publ. 19. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 24 p. [8287] 41. Taylor, R. F. 1932. The successional trend and its relation to second-growth forests in southeastern Alaska. Ecology. 13(4): 381-391. [10007] 42. U.S. Department of Agriculture, Forest Service, Alaska Region. [n.d.]. Preliminary forest plant associations of the Stikine Area, Tongass National Forest. R10-TP-72. Portland, OR. 126 p. [19016] 43. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573] 44. Van Dersal, William R. 1938. Native woody plants of the United States, their erosion-control and wildlife values. Washington, DC: U.S. Department of Agriculture. 362 p. [4240] 45. Ver Hoef, Jay M.; Neiland, Bonita J.; Glenn-Lewin, David C. 1988. Vegetation gradient analysis of two sites in southeast Alaska. Northwest Science. 62(4): 171-180. [19175] 46. Viereck, L. A.; Dyrness, C. T.; Batten, A. R.; Wenzlick, K. J. 1992. The Alaska vegetation classification. Gen. Tech. Rep. PNW-GTR-286. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 278 p. [2431] 47. Vitt, Dale H.; Horton, Diana G.; Slack, Nancy G.; Malmer, Nils. 1990. Sphagnum-dominated peatlands of the hyperoceanic British Columbia coast: patterns in surface water chemistry and vegetation. Canadian Journal of Forestry Research. 20: 696-711. [11739] 48. Vogl, Richard J.; Armstrong, Wayne P.; White, Keith L.; Cole, Kenneth L. 1977. The closed-cone pines and cypress. In: Barbour, Michael G.; Major, Jack, eds. Terrestrial vegetation of California. New York: John Wiley and Sons: 295-358. [7219] 49. Wheeler, Nicholas C.; Critchfield, William B. 1985. The distribution and botanical characteristics of lodgepole pine: biogeographical and management implications. In: Baumgartner, David M.; Krebill, Richard G.; Arnott, James T.; Weetman, Gordon F., compilers and editors. Lodgepole pine: The species and its management: Symposium proceedings; 1984 May 8-10; Spokane, WA; 1984 May 14-16; Vancouver, BC. Pullman, WA: Washington State University, Cooperative Extension: 1-13. [9435] 50. Owens, John N.; Molder, Marje. 1975. Development of long-shoot terminal buds of Pinus contorta spp. contorta. In: Baumgartner, David M., ed. Management of lodgepole pine ecosystems: Symposium proceedings; 1973 October 9-11; Pullman, WA. Vol. 1. Pullman, WA: Washington State University, Cooperative Extension Service: 86-104. [7822]

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