Index of Species Information
SPECIES: Liquidambar styraciflua
Introductory
SPECIES: Liquidambar styraciflua
AUTHORSHIP AND CITATION :
Coladonato, Milo. 1992. Liquidambar styraciflua. In: Fire Effects Information System, [Online].
U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station,
Fire Sciences Laboratory (Producer). Available:
https://www.fs.usda.gov/database/feis/plants/tree/liqsty/all.html [].
ABBREVIATION :
LIQSTY
SYNONYMS :
NO-ENTRY
SCS PLANT CODE :
LIST2
COMMON NAMES :
sweetgum
redgum
sapgum
star-leaf gum
blisted
satin-walnut
white gum
alligator-tree
opossum-tree
gum-wood
copalm balsam
TAXONOMY :
The currently accepted scientific name for sweetgum is Liquidambar
styraciflua L. [30]. Two forms of sweetgum are recognized in
horticulture. The round-lobed American sweetgum, L. styraciflua forma
rotundiloba Rehd., has three to five short, rounded lobes on the leaves.
Weeping American sweetgum, L. styraciflua forma pendula Rehd., has
pendulous branches forming an almost columnar head [44,45]. There are no
recognized subspecies or varieties.
LIFE FORM :
Tree
FEDERAL LEGAL STATUS :
No special status
OTHER STATUS :
NO-ENTRY
DISTRIBUTION AND OCCURRENCE
SPECIES: Liquidambar styraciflua
GENERAL DISTRIBUTION :
Sweetgum grows from Connecticut southward throughout the East to central
Florida and eastern Texas. It is found as far west as Missouri,
Arkansas, and Oklahoma and as far north as southern Illinois. It also
grows in scattered locations in northeastern and central Mexico,
Guatemala, Belize, El Salvador, Honduras, and Nicaragua [14,24,42]. It
is cultivated in Hawaii [50].
ECOSYSTEMS :
FRES12 Longleaf - slash pine
FRES13 Loblolly - shortleaf pine
FRES14 Oak - pine
FRES15 Oak - hickory
FRES16 Oak - gum - cypress
STATES :
AL AR DE FL GA HI IL IN KY LA
MD MS NJ OH OK PA SC TN TX VA
WV MEXICO
BLM PHYSIOGRAPHIC REGIONS :
NO-ENTRY
KUCHLER PLANT ASSOCIATIONS :
K097 Southeastern spruce - fir forest
K100 Oak - hickory forest
K102 Beech - maple forest
K103 Mixed mesophytic forest
K104 Appalachian oak forest
SAF COVER TYPES :
44 Chestnut oak
51 White pine - chestnut oak
52 White oak - black oak - northern red oak
57 Yellow-poplar
61 River birch - sycamore
62 Silver maple - American elm
64 Sassafras - persimmon
65 Pin oak - sweetgum
70 Longleaf pine
74 Cabbage palmetto
75 Shortleaf pine
76 Shortleaf pine - oak
78 Virginia pine - oak
79 Virginia pine
80 Loblolly pine - shortleaf pine
81 Loblolly pine
82 Loblolly pine - hardwood
83 Longleaf pine - slash pine
84 Slash pine
87 Sweetgum - yellow-poplar
88 Willow oak - water oak - diamondleaf oak
89 Live oak
91 Swamp chestnut - oak - cherrybark oak
92 Sweetgum - willow oak
93 Sugarberry - American elm - green ash
94 Sycamore - sweetgum - American elm
96 Overcup oak - water hickory
98 Pond pine
102 Baldcypress - tupelo
103 Water tupelo - swamp tupelo
104 Sweetbay - swamp tupelo - redbay
110 Black oak
SRM (RANGELAND) COVER TYPES :
NO-ENTRY
HABITAT TYPES AND PLANT COMMUNITIES :
NO-ENTRY
MANAGEMENT CONSIDERATIONS
SPECIES: Liquidambar styraciflua
WOOD PRODUCTS VALUE :
Sweetgum is primarily used for lumber, veneer, and plywood. The lumber
is used to make boxes, crates, furniture, interior trim, and millwork.
The veneer is used primarily for crates, baskets, and interior woodwork.
Sweetgum is also used for crossties and fuel, and small amounts go into
fencing, excelsior, and pulpwood [37,42].
IMPORTANCE TO LIVESTOCK AND WILDLIFE :
Sweetgum has moderate value as a winter browse [5]. In the Oconee
National Forest of Georgia, sweetgum was lightly to moderately browsed
by white-tailed deer during the fall and winter [19]. The seeds are
eaten by birds, squirrels, and chipmunks [33].
PALATABILITY :
NO-ENTRY
NUTRITIONAL VALUE :
Sweetgum is relatively high in protein and caloric content. Mean
nutrient values for sweetgum on unburned plots on the Siecke
State Forest, Texas, varied seasonally as follows [27]:
crude N-free
protein fat fiber extract ash phosphorus calcium
Spring 10.76 2.78 9.08 58.49 3.84 6.13 0.63
Summer 7.00 2.78 12.09 59.39 3.73 0.07 0.86
Fall 5.74 3.09 11.08 59.72 5.33 0.06 1.28
Winter 4.42 2.51 20.23 54.64 3.21 0.06 1.70
COVER VALUE :
Sweetgum snags are used as breeding sites for a variety of birds and
mammals [13].
VALUE FOR REHABILITATION OF DISTURBED SITES :
Sweetgum stem cuttings have been successfully planted for streambank
protection and reclamation of sites disturbed by coal strip mining
[29,46].
Sweetgum growth and survival was good when planted on favorable sites
but decreased when seedlings were planted concurrently with ground cover
or in previously established cover of grasses and legumes on mined sites
in southeastern Indiana [3,8].
OTHER USES AND VALUES :
Medicinally, sweetgum is known as "copalm balsam" and the resinous gum
is used extensively in Mexico and Europe as a substitute for storax.
Various ointments and syrups are prepared from the resinous gum and are
used in the treatment of dysentery and diarrhea. The gum is sometimes
chewed by children, and it is also used as a perfuming agent in soap
[45].
The beautiful red and yellow color variations of sweetgum's autumn
foliage make it highly prized as an ornamental [33,45].
OTHER MANAGEMENT CONSIDERATIONS :
Sweetgum's ability to sprout quickly and persistently makes it one of the
most serious competitors of pine seedlings in southeastern forests.
Silvicultural practices have called for the control of sweetgum in areas
where it competes heavily with pine seedlings [49]. Basal applications
of Garlon 4 top-killed 81 percent of 2 inch (5 cm) d.b.h or smaller
stems [35,36].
BOTANICAL AND ECOLOGICAL CHARACTERISTICS
SPECIES: Liquidambar styraciflua
GENERAL BOTANICAL CHARACTERISTICS :
Sweetgum is a large, native, long-lived, deciduous tree that reaches
heights of 50 to 150 feet (15-45 m) at maturity [6,14]. It is easily
recognizable by the long-petioled, star-shaped leaves which have five
long-pointed, saw-toothed lobes. The brown bark is deeply furrowed into
narrow scaley plates or ridges. Young sweetgum trees have long conical
crowns, while mature trees have crowns that are round and spreading.
Sweetgum is monoecious with the male flowers in several clusters and the
female flowers hanging at the end of the same stalk. The ball-shaped
fruits contain many individual seed-bearing sections, and persist
throughout the winter [16,18].
RAUNKIAER LIFE FORM :
Phanerophyte
REGENERATION PROCESSES :
Seed production and dissemination: Sweetgum produces an abundance of
lightweight seed. The tree begins to produce seed when 20 to 30 years
old, and crops remain abundant for 150 years. Fair seed crops are
produced each year, with bumper crops every 2 to 3 years [2,24]. Under
conditions of full sunlight and rich moist soil, each fruit may average
as many as 50 sound seeds. Seed is primarily dispersed by wind; the
maximum dispersal distance recorded was 600 feet (183 m) but
ordinarily 96 percent of the seed fall within 200 feet (61 m) of the
point of release [24,38].
Seedling development: Sod is not a serious hindrance to seed
germination; however, when additional sweetgum production is desired in
partially cutover stands, exposed mineral soil and abundant direct
sunlight are necessary [4,22]. Root development varies with the growing
site. A deep taproot and numerous horizontal rootlets usually develop
early, but in wet areas the root system is shallow and wide spreading,
with little or no taproot [25,39]. On an abandoned field adjacent to a
swamp in Maryland, 5-year-old seedlings averaged 8.7 feet (2.6 m) in
height [24]. On favorable sites in the lower Mississippi Valley,
seedlings grow as much as 2 feet (0.6 m) during the first year [24,49].
Vegetative reproduction: Sweetgum is capable of sprouting until it is
approximately 50 years old. Although sweetgum seedlings reach a height
of 4.5 feet (1.4 m) in 3 to 5 years, sprouts often reach this height in
one growing season. Ten-year old sprouts frequently have the same size
and appearance as 18- to 20-year-old seedlings in the same stand
[23,49].
SITE CHARACTERISTICS :
Sweetgum is very tolerant of different soils and sites but grows best on
the rich, moist, alluvial clay and loamy soils of river bottoms [28].
Throughout the Piedmont Plateau, sweetgum shows good growth on river and
stream bottoms and shows considerable potential on many upland sites
[24,34].
Common tree associates of sweetgum include spruce pine (Pinus glabra),
Virginia pine (P. virginiana), red maple (Acer rubrum), box elder (A.
negundo), pignut, shellbark, shagbark, and mockernut hickories (Carya
glabra, C. laciniosa, C. ovata, C. tomentosa), and sugarberry (Celtis
laevigata). Common understory associates include dogwood (Cornus spp.),
alder (Alnus spp.), and eastern redbud (Cercis canadensis) [1,10,24].
SUCCESSIONAL STATUS :
Sweetgum is classified as shade intolerant [7]. In pure stands on
bottomland sites, young sweetgum is able to endure some shade and
crowding. With increase in age the tree becomes less tolerant of
competition. Following natural decrease in the canopy, enough sunlight
reaches the ground to permit an understory stand to develop [12,24].
Although sweetgum is an early invader, it seldom becomes a dominant
species [20,31].
SEASONAL DEVELOPMENT :
Sweetgum flowers appear from March to May, depending on latitude and
weather. The fruit ripens from September to November; the fruit often
persists through the entire winter [6,24].
FIRE ECOLOGY
SPECIES: Liquidambar styraciflua
FIRE ECOLOGY OR ADAPTATIONS :
Fire is one of the major agents of damage to sweetgum. Its relatively
thin bark make it highly susceptible to fire [21]. Following top-kill
by fire, sweetgum sprouts from the stump or root crown [41,48].
FIRE REGIMES :
Find fire regime information for the plant communities in which this
species may occur by entering the species name in the FEIS home page under
"Find Fire Regimes".
POSTFIRE REGENERATION STRATEGY :
survivor species; on-site surviving root crown or caudex
off-site colonizer; seed carried by wind; postfire years 1 and 2
FIRE EFFECTS
SPECIES: Liquidambar styraciflua
IMMEDIATE FIRE EFFECT ON PLANT :
Fire typically top-kills sweetgum. Hot summer fires may deplete
carbohydrate reserves and eventually kill the tree [41,48].
DISCUSSION AND QUALIFICATION OF FIRE EFFECT :
Fire scars on living trees provide entry points for insects and
diseases. As long as the sapwood is not killed by fire, basal wounds
are often covered with a gum exudation that protects them. After
repeated fires, however, a tree is apt to have some sapwood killed and
fungi and insects may become established [24,47].
PLANT RESPONSE TO FIRE :
Sweetgum generally sprouts prolifically when top-killed by fire.
Repeated annual summer burns, however, will eventually deplete
carbohydrate reserves and kill the plant [41,48].
DISCUSSION AND QUALIFICATION OF PLANT RESPONSE :
NO-ENTRY
FIRE MANAGEMENT CONSIDERATIONS :
Fire has been demonstrated to be a good management tool for controlling
sweetgum. In the Coastal Plain of South Carolina, five consecutive
summer fires killed 85 percent or more of the root stalks of sweetgum.
Winter fires did not kill appreciable numbers of root stalks but did
top-kill most sweetgum 2 inches (5 cm) or less d.b.h. [11].
REFERENCES
SPECIES: Liquidambar styraciflua
REFERENCES :
1. Abernethy, Y.; Turner, R. E. 1987. US forested wetlands: 1940-1980:
Field-data surveys document changes and can guide national resource
management. BioScience. 37(10): 721-727. [10575]
2. Allen, James A.; Kennedy, Harvey E., Jr. 1989. Bottomland hardwood
reforestation in the lower Mississippi Valley. Slidell, LA: U.S.
Department of the Interior, Fish and Wildlife Service, National Wetlands
Research Center; Stoneville, MS: U.S. Department of Agriculture, Forest
Service, Southern Forest Experimental Station. 28 p. [15293]
3. Andersen, C. P.; Bussler, B. H.; Chaney, W. R.; [and others]. 1989.
Concurrent establishment of ground cover and hardwood trees on reclaimed
mined land and unmined reference sites. Forest Ecology and Management.
28: 81-99. [10920]
4. Belcher, Earl W., Jr.; Hitt, Robert G. 1965. Eastern Tree Seed
Laboratory: 12th annual report, fiscal year 1965. Macon, GA: Eastern
Tree Seed Laboratory. 66 p. In cooperation with: Region 8 and the
Southeastern Forest Experiment Station, U.S. Forest Service; Georgia
Forestry Commission and Georgia Forest Research Council. [6522]
5. Blair, Robert M.; Short, Henry L.; Epps, E. A., Jr. 1977. Seasonal
nutrient yield and digestibility of deer forage from a young pine
plantation. Journal of Wildlife Management. 41(4): 667-676. [16963]
6. Bonner, F. T. 1974. Liquidambar styraciflua L. sweetgum. In:
Schopmeyer, C. S., ed. Seeds of woody plants in the United States.
Agriculture Handbook No. 450. Washington, DC: U.S. Department of
Agriculture, Forest Service: 505-507. [7695]
7. Borthwick, H. A. 1957. Light effects on tree growth and seed
germination. A symposium on forest tree physiology; 1957 June 13-14;
Wooster, OH. In: The Ohio Journal of Science; 57(6): 357-364. [10162]
8. Brothers, Timothy S. 1988. Indiana surface-mine forests: historical
development and composition of a human-created vegetation complex.
Southeastern Geographer. 28(1): 19-33. [8787]
9. Brown, Arthur A.; Davis, Kenneth P. 1973. Forest fire control and use.
2nd ed. New York: McGraw-Hill. 686 p. [15993]
10. Carter, Katherine K.; Snow, Albert G., Jr. 1990. Pinus virginiana Mill.
Virginia pine. In: Burns, Russell M.; Honkala, Barbara H., technical
coordinators. Silvics of North America. Volume 1. Conifers. Agric.
Handb. 654. Washington, DC: U.S. Department of Agriculture, Forest
Service: 513-519. [13411]
11. Chen, Ming-Yih; Hodgkins, Earl J.; Watson, W. J. 1975. Prescribed
burning for improving pine production and wildlife habitat in the hilly
coastal plain of Alabama. Bull. No. 473. Auburn, AL: Auburn University
Agricultural Experiment Station. 19 p. [9909]
12. Clatterbuck, Wayne K. 1989. Even-aged mixtures of cherrybark oak and
loblolly pine in southwestern Arkansas. In: Waldrop, Thomas A., ed.
Proceedings of pine-hardwood mixtures: a symposium on management and
ecology of the type; 1989 April 18-19; Atlanta, GA. Gen. Tech. Rep.
SE-58. Asheville, SC: U.S. Department of Agriculture, Forest Service,
Southeastern Forest Experiment Station: 123-127. [10268]
13. Dickson, James G.; Conner, Richard N.; Williamson, J. Howard. 1983. Snag
retention increases bird use of a clear-cut. Journal of Wildlife
Management. 47(3): 799-804. [13855]
14. Duncan, Wilbur H.; Duncan, Marion B. 1987. The Smithsonian guide to
seaside plants of the Gulf and Atlantic Coasts from Louisiana to
Massachusetts, exclusive of lower peninsular Florida. Washington, DC:
Smithsonian Institution Press. 409 p. [12906]
15. Eyre, F. H., ed. 1980. Forest cover types of the United States and
Canada. Washington, DC: Society of American Foresters. 148 p. [905]
16. Fernald, Merritt Lyndon. 1950. Gray's manual of botany. [Corrections
supplied by R. C. Rollins]. Portland, OR: Dioscorides Press. 1632 p.
(Dudley, Theodore R., gen. ed.; Biosystematics, Floristic & Phylogeny
Series; vol. 2). [14935]
17. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others].
1977. Vegetation and environmental features of forest and range
ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of
Agriculture, Forest Service. 68 p. [998]
18. Godfrey, Robert K. 1988. Trees, shrubs, and woody vines of northern
Florida and adjacent Georgia and Alabama. Athens, GA: The University of
Georgia Press. 734 p. [10239]
19. Harlow, Richard F.; Shrauder, Paul A.; Seehorn, Monte E. 1975. Deer
browse resources of the Oconee National Forest. Res. Pap. SE-137.
Asheville, NC: U.S. Department of Agriculture, Forest Service,
Southeastern Forest Experiment Station. 16 p. [14602]
20. Hartnett, David C.; Krofta, Douglas M. 1989. Fifty-five years of
post-fire succession in a southern mixed hardwood forest. Bulletin of
the Torrey Botanical Club. 116(2): 107-113. [9153]
21. Hodgkins, Earl J. 1958. Effects of fire on undergrowth vegetation in
upland southern pine forests. Ecology. 39(1): 36-46. [7632]
22. Hunt, Ron; Cleveland, Glenn. 1978. Cultural treatments affect growth,
volume, and survival of sweetgum, sycamore, and loblolly pine. Southern
Journal of Applied Forestry. 2(2): 55-59. [10633]
23. Kaufert, F. H. 1933. Fire and decay injury in the Southern bottomland
hardwoods. Journal of Forestry. 31: 64-67. [2694]
24. Kormanik, Paul P. 1990. Liquidambar styraciflua L. sweetgum. In: Burns,
Russell M.; Honkala, Barbara H., technical coordinators. Silvics of
North America. Vol. 2. Hardwoods. Agric. Handb. 654. Washington, DC:
U.S. Department of Agriculture, Forest Service: 400-405. [17401]
25. Kormanik, P. P.; Ruehle, J. L. 1987. Lateral root development may define
nursery seedling quality. In: Phillips, Douglas R., compiler.
Proceedings. 4th biennial southern silvicultural research conference;
1986 November 4-6; Atlanta, GA. Gen. Tech. Rep. SE-42. Asheville, NC:
U.S. Department of Agriculture, Forest Service, Southeastern Forest
Experiment Station: 225-229. [4199]
26. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation
of the conterminous United States. Special Publication No. 36. New York:
American Geographical Society. 77 p. [1384]
27. Lay, Daniel W. 1957. Browse quality and the effects of prescribed
burning in southern pine forests. Journal of Forestry. 55: 342-347.
[7633]
28. Lea, Russ; Frederick, D. J. 1990. Bottomland hardwood restoration in the
southeastern United States. In: Hughes, H. Glenn; Bonnicksen, Thomas M.,
eds. Restoration `89: the new management challenge: Proceedings, 1st
annual meeting of the Society for Ecological Restoration; 1989 January
16-20; Oakland, CA. Madison, WI: The University of Wisconsin Arboretum,
Society for Ecological Restoration: 292-300. [14706]
29. Limstrom, G. A.; Merz, R. W. 1949. Rehabilitation of lands stripped for
coal in Ohio. Tech. Pap. No. 113. Columbus, OH: The Ohio Reclamation
Association. 41 p. In cooperation with: U.S. Department of Agriculture,
Forest Service, Central States Forest Experiment Station. [4427]
30. Little, Elbert L., Jr. 1979. Checklist of United States trees (native
and naturalized). Agric. Handb. 541. Washington, DC: U.S. Department of
Agriculture, Forest Service. 375 p. [2952]
31. Lotti, Thomas; Klawitter, Ralph A.; LeGrande, W. P. 1960. Prescribed
burning for understory control in loblolly pine stands of the coastal
plain. Station Pap. No. 116. Asheville, NC: U.S. Department of
Agriculture, Forest Service, Southeastern Forest Experiment Station. 19
p. [15417]
32. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession
following large northern Rocky Mountain wildfires. In: Proceedings, Tall
Timbers fire ecology conference and Intermountain Fire Research Council
fire and land management symposium; 1974 October 8-10; Missoula, MT. No.
14. Tallahassee, FL: Tall Timbers Research Station: 355-373. [1496]
33. Martin, Alexander C.; Zim, Herbert S.; Nelson, Arnold L. 1951. American
wildlife and plants. New York: McGraw-Hill Book Company, Inc. 500 p.
[4021]
34. McGarity, R. W. 1979. Young sweetgum responds to early merchantable
thinning. Southern Journal of Applied Forestry. 3(4): 157-160. [10617]
35. McLemore, B. F.; Cain, M. D. 1988. A test of basal sprays for
controlling hardwood brush and trees. In: Environmental legislation and
its effect on weed science: Proceedings, 41st annual meeting Southern
Weed Science Soc; 1988 January 18-20; Tulsa, OK. Volume 41. [Place of
publication unknown]. [Publisher unknown]. 180-186. [12154]
36. Miller, James H. 1990. Streamline basal application of herbicide for
small-stem hardwood control. Southern Journal of Applied Forestry.
14(4): 161-165. [13538]
37. Millers, Imants; Shriner, David S.; Rizzo, David. 1989. History of
hardwood decline in the eastern United States. Gen. Tech. Rep. NE-126.
Bromall, PA: U.S. Department of Agriculture, Forest Service,
Northeastern Forest Experiment Station. 75 p. [10925]
38. Paton, Robert R.; Secrest, Edmund; Ezri, Harold A. 1944. Ohio forest
plantings. Bull. 647. Wooster, OH: Ohio Agricultural Experiment Station.
77 p. [6974]
39. Powell, Susan W.; Day, Frank P., Jr. 1991. Root production in four
communities in the Great Dismal Swamp. American Journal of Botany.
78(2): 288-297. [14595]
40. Raunkiaer, C. 1934. The life forms of plants and statistical plant
geography. Oxford: Clarendon Press. 632 p. [2843]
41. Riebold, R. J. 1955. Summer burns for hardwoods control in loblolly
pine. Fire Control Notes. 16(1): 34-35. [11602]
42. U.S. Department of Agriculture, Forest Service. 1956. Wood...colors and
kinds. Agric. Handb. 101. Washington, DC. 36 p. [16294]
43. U.S. Department of Agriculture, Soil Conservation Service. 1982.
National list of scientific plant names. Vol. 1. List of plant names.
SCS-TP-159. Washington, DC. 416 p. [11573]
44. Van Dersal, William R. 1938. Native woody plants of the United States,
their erosion-control and wildlife values. Washington, DC: U.S.
Department of Agriculture. 362 p. [4240]
45. Vines, Robert A. 1960. Trees, shrubs, and woody vines of the Southwest.
Austin, TX: University of Texas Press. 1104 p. [7707]
46. Vogel, Willis G. 1981. A guide for revegetating coal minespoils in the
eastern United States. Gen. Tech. Rep. NE-68. Broomall, PA: U.S.
Department of Agriculture, Forest Service, Northeastern Forest
Experiment Station. 190 p. [15577]
47. Waldrop, Thomas A.; Lloyd, F. Thomas. 1991. Forty years of prescribed
burning on the Santee fire plots: effects on overstory and midstory
vegetation. In: Nodvin, Stephen C.; Waldrop, Thomas A., eds. Fire and
the environment: ecological and cultural perspectives: Proceedings of an
international symposium; 1990 March 20-24; Knoxville, TN. Gen. Tech.
Rep. SE-69. Asheville, NC: U.S. Department of Agriculture, Forest
Service, Southeastern Forest Experiment Station: 45-50. [16632]
48. Waldrop, Thomas A.; Van Lear, David H.; Lloyd, F. Thomas; Harms, William
R. 1987. Long-term studies of prescribed burning in loblolly pine
forests of the Southeastern Coastal Plain. Gen. Tech. Rep. SE-45.
Asheville, NC: U.S. Department of Agriculture, Forest Service,
Southeastern Forest Experiment Station. 23 p. [11596]
49. Wenger, Karl F. 1953. The sprouting of sweetgum in relation to season of
cutting and carbohydrate content. Plant Physiology. 28: 35-49. [13819]
50. St. John, Harold. 1973. List and summary of the flowering plants in the
Hawaiian islands. Hong Kong: Cathay Press Limited. 519 p. [25354]
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